:: wikimiki.org ::

Eye

Eye

: This article refers to the sight organ. See Eye (disambiguation) for other usages An eye is an organ that detects light. Different kinds of light-sensitive organs are found in a variety of creatures. The simplest eyes do nothing but detect whether the surroundings are light or dark. More complex eyes are used to provide the sense of vision. Many complex organisms including some mammals, birds, reptiles and fish have two eyes which may be placed on the same plane to be interpreted as a single three-dimensional "image" (binocular vision), as in humans; or on different planes producing two separate "images" (monocular vision), such as in rabbits and chameleons.
Varieties of eyes
chameleon chameleon]] In most vertebrates and some mollusks the eye works by allowing light to enter it and project onto a light-sensitive panel of cells known as the retina at the rear of the eye, where the light is detected and converted into electrical signals, which are then transmitted to the brain via the optic nerve. Such eyes are typically roughly spherical, filled with a transparent gel-like substance called the vitreous humour, with a focusing lens and often an iris which regulates the intensity of the light that enters the eye. The eyes of cephalopods, fish, amphibians, and snakes usually have fixed lens shapes, and focusing vision is achieved by telescoping the lens (similar to how a camera focuses). Compound eyes are found among the arthropods and are composed of many simple facets which give a pixelated image (not multiple images as is often believed). Each sensor has its own lens and photosensitive cell(s). Some eyes have up to 28,000 such sensors, which are arranged hexagonally, and which can give a full 360 degree field of vision. Compound eyes are very sensitive to motion. Some arthropods (many Strepsiptera) have compound eye composed of a few facets each with a retina capable of creating an image, which does provide muliple image vision. With each eye viewing a different angle, a fused image from all the eyes is produced in the brain providing a very wide angle high resolution image. Trilobites, which are now extinct, had unique compound eyes. They used clear calcite crystals to form the lenses of their eyes. In this, they differ from most other arthropods, which have soft eyes. The number of lenses in such an eye varied, however: some trilobites had only one, and some had thousands of lenses in one eye. Some of the simplest eyes, called ocelli, can be found in animals like snails, who can not actually "see" in the common sense. They do have photosensitive cells, but no lens and no other means of projecting an image onto these cells. They can distinguish between light and dark (day and night), but no more. This enables snails to keep out of direct sunlight. Jumping spiders have simple eyes that are so large, supported by an array of other smaller eyes, that they can get enough visual inputs to hunt and pounce on their prey. Some insect larvae like caterpillars have a different type of single eye (stemmata) which gives a rough image.
Evolution of eyes
How a complex structure like the projecting eye could have evolved is often said to be a difficult question for the theory of evolution. Darwin famously treated the subject of eye evolution in his Origin of Species: :To suppose that the eye, with all its inimitable contrivances for adjusting the focus to different distances, for admitting different amounts of light, and for the correction of spherical and chromatic aberration, could have been formed by natural selection, seems, I freely confess, absurd in the highest possible degree. Yet reason tells me, that if numerous gradations from a perfect and complex eye to one very imperfect and simple, each grade being useful to its possessor, can be shown to exist; if further, the eye does vary ever so slightly, and the variations be inherited, which is certainly the case; and if any variation or modification in the organ be ever useful to an animal under changing conditions of life, then the difficulty of believing that a perfect and complex eye could be formed by natural selection, though insuperable by our imagination, can hardly be considered real. Despite the precision and complexity of the eye, computer models of eye evolution, developed by Dan-Erik Nilsson and Susanne Pelger, demonstrated that a primitive optical sense organ could evolve into a complex human-like eye within a reasonable period (less than a million years) simply through small mutations and natural selection. Eyes in various animals show adaption to their requirements. For example, birds of prey have much greater visual acuity than humans and some, like diurnal birds of prey, can see ultraviolet light. The different forms of eye in, for example, vertebrates and mollusks are often cited as examples of parallel evolution, suggesting that the development of eyes through evolution might not be so improbable as it might seem. However, the development of the eye is considered to be monophyletic; that is, all modern eyes, varied as they are, have their origins in a proto-eye believed to have evolved some 540 million years ago (Mya).
Anatomy
monophyletic monophyletic The structure of the mammalian eye owes itself completely to the task of focusing light onto the retina. All of the individual components through which light travels within the eye before reaching the retina are transparent, minimising dimming of the light. The cornea and lens help to converge light rays to focus onto the retina. This light causes chemical changes in the photosensitive cells of the retina, the products of which trigger nerve impulses which travel to the brain. Light enters the eye from an external medium such as air or water, passes through the cornea, and into the first of two humours, the aqueous humour. Most of the light refraction occurs at the cornea which has a fixed curvature. The first humour is a clear mass which connects the cornea with the lens of the eye, helps maintain the convex shape of the cornea (necessary to the convergence of light at the lens) and provides the corneal endothelium with nutrients. The iris, between the lens and the first humour, is a coloured ring of muscle fibres. Light must first pass though the centre of the iris, the pupil. The size of the pupil is actively adjusted by the circular and radial muscles to maintain a relatively constant level of light entering the eye. Too much light being let in could damage the retina, too little light would be blinding. The lens, behind the iris, is a convex, springy disk which focuses light, through the second humour, onto the retina. To clearly see an object far away, the circularly arranged ciliary muscles will pull on the lens, flattening it. Without muscles pulling on it, the lens will spring back into a thicker, more convex, form. Humans gradually lose this flexibility with age, resulting in the inability to focus on nearby objects, which is known as presbyopia. There are other refraction errors arising from the shape of the cornea and lens, and from the length of the eyeball. These include myopia, hyperopia, and astigmatism. On the other side of the lens is the second humour, the vitreous humour, which is bounded on all sides: by the lens, ciliary body, suspensory ligaments and by the retina. It lets light through without refraction, helps maintain the shape of the eye and suspends the delicate lens. Wrapped around these tissues are three layers of tissue surrounding the vitreous humour. The outermost is the sclera which gives the eye most of its white colour. It consists of fibrin connective tissue and both protects the inner components of the eye and maintains its shape. On the inner side of the sclera is the choroid, which contains blood vessels that supply the retinal cells with necessary oxygen and removes the waste products of respiration. Within the eye, only the sclera and ciliary muscles contain blood vessels. The choroid gives the inner eye a dark colour, which prevents disruptive reflections within the eye. The inner most layer of the eye is the retina, containing of the photosensitive rod and cone cells, and neurons. To maximise vision and light absorption, the retina is a relatively smooth (but curved) layer. It does have two points at which it is different; the fovea and blind spot. The fovea is a dip in the retina directly opposite the lens, which is densely packed with cone cells. It is largely responsible for colour vision in humans, and enables high acuity, such as is necessary in reading. The blind spot is a point on the retina where the optic nerve pierces the retina to connect to the nerve cells on its inside. No photosensitive cells exist at this point, it is thus "blind". In some animals, the retina contains a reflective layer (the tapetum lucidum) which increases the amount of light each photosensitive cell perceives, allowing the animal to see better under low light conditions.
Other articles regarding eye anatomy
Aqueous humour, Anterior chamber, Blind spot, Canal of Schlemm, Ciliary body, Ciliary muscle, Cornea, Conjunctiva, Choroid, Fovea, Iris, Lens, Macula, Optic disc, Optic nerve, Ora serrata, Posterior chamber, Pupil, Retina, Sclera, Suspensory ligament, Tapetum lucidum, Trabecular meshwork, Vitreous humour, Zonular fibers.
Cytology
The retina contains two forms of photosensitive cells - rods and cones. Though structurally and metabolically similar, their function is quite different, though they are equally important to vision. Rod cells are highly sensitive to light allowing them to respond in dim light and dark conditions. These are the cells which allow humans and other animals to see by moonlight, or with very little available light (as in a dark room). However, they do not distinguish between colours, and have low visual acuity (a measure of detail). This is why the darker conditions become, the less colour objects seem to have. Cone cells, conversely, need high light intensities to respond and have high visual acuity. Different cone cells respond to different colours (wavelengths) of light, which allows an organism to see colour. The differences are useful; apart from enabling sight in both dim and light conditions, humans have given them further application. The fovea, directly behind the lens, consists of mostly densely-packed cone cells. This gives humans a highly detailed central vision, allowing reading, bird watching, or any other task which primarily requires looking at things. Its requirement for high intensity light does cause problems for astronomers, as they cannot see dim stars, or other objects, using central vision because the light from these is not enough to stimulate cone cells. Because cone cells are all that exist directly in the fovea, astronomers have to look at stars through the "corner of their eyes" where rods also exist, and where the light is sufficient to stimulate cells, allowing the individual to observe distant stars. Rods and cones are both photosensitive, but respond differently to different frequencies of light. They both contain different pigmented photoreceptor proteins. Rod cells contain the protein rhodopsin and cone cells contain different proteins for each colour-range. The process through which these proteins go is quite similar - upon being subjected to electromagnetic radiation of a particular wavelength and intensity (ie. a colour visible light) the protein breaks down into two constituent products. Rhodopsin, of rods, breaks down into opsin and retinal; iodopsin of cones breaks down into photopsin and retinal. The opsin in both opens ion channels on the cell membrane which leads to the generation of an action potential (an impulse which will eventually get to the visual cortex in the brain). This is the reason why cones and rods enable organisms to see in dark and light conditions - each of the photoreceptor proteins requires a different light intensity to break down into the constituent products. Further, synaptic convergence means that several rod cells are connected to a single bipolar cell, which then connects to a single ganglion cell and information is relayed to the visual cortex. Whereas, a single cone cell is connected to a single bipolar cell. Thus, action potentials from rods share neurons, where those from cones are given their own. This results in the high visual acuity, or the high ability to distinguish between detail, of cone cells and not rods. If a ray of light were to reach just one rod cell this may not be enough to stimulate an action potential. Because several "converge" onto a bipolar cell, enough transmitter molecules reach the synapse of the bipolar cell to attain the threshold level to generate an action potential. Furthermore, colour is distinguishable when breaking down the iodopsin of cone cells because there are three forms of this protein. One form is broken down by the particular EM wavelength that is red light, another green light, and lastly blue light. In simple terms, this allows human beings to see red, green and blue light. If all three forms of cones are stimulated equally, then white is seen. If none are stimulated, black is seen. Most of the time however, the three forms are stimulated to different extents - resulting in different colours being seen. If, for example, the red and green cones are stimulated to the same extent, and no blue cones are stimulated, yellow is seen. For this reason red, green and blue are called primary colours and the products of mixing two secondary colours. The secondary colours can be further complimented with primary colours to see tertiary colours.
Acuity
Visual acuity can be measured with several different metrics. Cycles per degree (CPD) measures how much an eye can differentiate one object from another in terms of degree angles. It is essentially no different from angular resolution. To measure CPD, first draw a series of black and white lines of equal width on a grid (similar to a bar code). Next, place the observer at a distance such that the sides of the grid appear one degree apart. If the grid is 1 meter away, then the grid should be about 8.7 millimeters wide. Finally, increase the number of lines and decrease the width of each line until the grid appears as a solid grey block. In one degree, a human would not be able to distinguish more than about 12 lines without the lines blurring together. So a human can resolve distances of about 0.73 millimeters at a distance of one meter. A horse can resolve about 14 CPD (0.62 mm at 1 m) and a rat can resolve about 1 CPD (8.7 mm at 1 m). A diopter is the unit of measure of focus.
Dynamic range
At any given instant, the retina can resolve a contrast ratio of around 100:1 (about 6 1/2 stops). As soon as your eye moves (saccades) it re-adjusts its exposure both chemically and by adjusting the iris. Hence, over time, a contrast ratio of about 1,000,000:1 (about 20 stops) can be resolved.
Adnexa and related parts

The orbit
In many species, the eyes are inset in the portion of the skull known as the orbits or eyesockets. This placement of the eyes helps to protect them from injury.
Eyebrows
In humans, the eyebrows redirect flowing substances (usually rainwater) away from the eye. Water in the eye can alter the refractive properties of the eye and blur vision. It can also wash away the tear fluid, and its beneficial effects, and can damage the cornea, due to osmotic differences between tear fluid and freshwater.
Eyelids
In many animals, including humans, eyelids wipe the eye and prevent the eyes from dehydration. They spread tear fluid on the eyes, which contains substances which help fight bacterial infection as part of the immune system. Some aquatic animals have a second eyelid in each eye which refracts the light and helps them see clearly both above water and below it. Most creatures will automatically react to a threat to its eyes (such as an object moving straight at the eye, or a bright light) by covering the eyes, and/or by turning the eyes away from the threat. Blinking the eyes is, of course, also a reflex.
Eyelashes
In many animals, including humans, eyelashes prevent fine particles from entering the eye. Fine particles can be bacteria, but also simple dust which can cause irritation of the eye, and lead to tears and subsequent blurred vision.
Eye movement
Animals with compound eyes have a wide field of vision, allowing them to look in many directions. To see more, they have to move their entire head or even body. The visual system in the brain is too slow to process that information if the images are slipping across the retina at more than a few degrees per second (Westheimer and McKee, 1954). Thus, for humans to be able to see while moving, the brain must compensate for the motion of the head by turning the eyes. Another complication for vision in frontal-eyed animals is the development of a small area of the retina with a very high visual acuity. This area is called the fovea, and covers about 2 degrees of visual angle in people. To get a clear view of the world, the brain must turn the eyes so that the image of the object of regard falls on the fovea. Eye movements are thus very important for visual perception, and any failure to make them correctly can lead to serious visual disabilities. To see a quick demonstration of this fact, try the following experiment: hold your hand up, about one foot (30 cm) in front of your nose. Keep your head still, and shake your hand from side to side, slowly at first, and then faster and faster. At first you will be able to see your fingers quite clearly. But as the frequency of shaking passes about one hertz, the fingers will become a blur. Now, keep your hand still, and shake your head (up and down or left and right). No matter how fast you shake your head, the image of your fingers remains clear. This demonstrates that the brain can move the eyes opposite to head motion much better than it can follow, or pursue, a hand movement. When your pursuit system fails to keep up with the moving hand, images slip on the retina and you see a blurred hand. Having two eyes is an added complication, because the brain must point both of them accurately enough that the object of regard falls on corresponding points of the two retinas; otherwise, double vison would occur. The movements of different body parts are controlled by striated muscles acting around joints. The movements of the eye are no exception, but they have special advantages not shared by skeletal muscles and joints, and so are considerably different.
Extraocular muscles
Each eye has six muscles that control its movements: the lateral rectus, the medial rectus, the inferior rectus, the superior rectus, the inferior oblique, and the superior oblique. When the muscles exert different tensions, a torque is exerted on the globe that causes it to turn. This is an almost pure rotation, with only about one millimeter of translation (Carpenter, 1988). Thus, the eye can be considered as undergoing rotations about a single joint in the center of the eye.
Rapid eye movement
Rapid eye movement typically refers to the stage during sleep during which the most vivid dreams occur. During this stage, the eyes move rapidly. It is not in itself a unique form of eye movement.
Saccades
Saccades are rapid refocussing actions of the eyes. Many animals are able to quickly look at a point in space (prompted by memory, peripheral vision or an audio cue) without actively looking at anything in between. The eyes simply jerk into a new position. Saccades move the eye at up to 900°/s in adult humans.
Microsaccades
Even when looking intently at a single spot, the eyes drift around. This ensures that individual photosensitive cells are continually stimulated in different degrees. Without changing input, these cells would otherwise stop generating output. Microsaccades move the eye no more than a total of 0.2° in adult humans.
Vestibulo-ocular reflex
Many animals can look at something while turning their heads. The eyes are automatically rotated to remain fixed on the object, directed by input from the organs of balance near the ears.
Smooth pursuit movement
The eyes can also follow a moving object around. This is less accurate than the vestibulo-ocular reflex as it requires the brain to process incoming visual information and supply feedback. Following an object moving at constant speed is relatively easy, though the eyes will often make saccadic jerks to keep up. The smooth pursuit movement can move the eye at up to 100°/s in adult humans.
Optokinetic reflex
The optokinetic reflex is a combination of a saccade and smooth pursuit movement. When, for example, looking out of the window in a moving train, the eyes can focus on a 'moving' tree for a short moment (through smooth pursuit), until the tree moves out of the field of vision. At this point, the optokinetic reflex kicks in, and moves the eye back to the point where it first saw the tree (through a saccade).
Vergence movement
feedback When a creature with binocular vision looks at an object, the eyes must rotate around a vertical axis so that the projection of the image is in the centre of the retina in both eyes. To look at an object closer by, the eyes rotate 'towards each other' (convergence), while for an object farther away they rotate 'away from eachother' (divergence). Exaggerated convergence is called cross eyed viewing (focussing on the nose for example) . When looking into the distance, or when 'staring into nothingness', the eyes neither converge nor diverge. Vergence movements are closely connected to accommodation of the eye. Under normal conditions, changing the focus of the eyes to look at an object at a different distance will automatically cause vergence and accommodation.
Accommodation
To see clearly, the lens will be pulled flatter or allowed to regain its thicker form.
Diseases, disorders, and age-related changes
There are many diseases and disorders that may affect the eyes. As the eye ages certain changes occur that can be attributed to solely the aging process. Most of these anatomic and physiologic processes follow a gradual decline. With aging, the quality of vision worsens due to reasons independent of aging eye diseases. While there are many changes of significance in the nondiseased eye, the most functionally important changes seem to be a reduction in pupil size and the loss of accommodation or focusing capability (presbyopia). The area of the pupil governs the amount of light that can reach the retina. The extent to which the pupil dilates also decreases with age. Because of the smaller pupil size, older eyes receive much less light at the retina. In comparison to younger people, it is as though older persons wear medium-density sunglasses in bright light and extremely dark glasses in dim light. Therefore, for any detailed visually guided tasks on which performance varies with illumination, older person requires extra lighting. With aging a prominent white ring develops in the periphery of the cornea- called arcus senilis. Aging causes laxity and downward shift of eyelid tissues and atrophy of the orbital fat. These changes contribute to the etiology of several eyelid disorders such as ectropion, entropion, dermatochalasis,and ptosis. The vitreous gel undergoes liquefaction (posterior vitreous detachment or PVD) and its opacities - visible as floaters gradually increase in number.
See also

- WikiSaurus:eye — the WikiSaurus list of synonyms and slang words for eyes in many languages
- Adaptation
- Binocular vision
- Corrective lens
- Crystallin
- Evil eye
- Eye color
- Eye contact
- Eye tracking
- Eyeglass prescription
- Macropsia
- Micropsia
- Nictitating membrane
- Ocular tremor
- Ophthalmology
- Optician
- Optometry
- Persistence of vision
- Phosphenes
- Snellen chart
- Staring contest
- Tears
- Visual perception
External links

- [http://www.djo.harvard.edu/ DJO | Digital Journal of Ophthalmology]
- [http://www.afb.org/eyeconditions.asp Glossary of Eye Conditions]
- [http://www.pbs.org/wgbh/evolution/library/01/1/l_011_01.html Evolution of the Eye]
- [http://www.eyetopics.com eye Topics]
- [http://webvision.med.utah.edu/anatomy.html Diagram of the eye]
- [http://webvision.med.utah.edu/ Webvision. The organisation of the retina and visual system.]
References

-
-
- [http://soma.npa.uiuc.edu/courses/bio303/Ch11b.html Internet lecture on eye types in animal kindom] # [http://www.agingeye.net/ AgingEye Times] Category:Visual system Category:Head and neck Category:Ophthalmology ms:Mata ja:目 zh-min-nan:Ba̍k-chiu

Eye (disambiguation)
The article Eye refers to the sight organ. Here are some other usages of the word "eye":
- A sewing needle has an eye to thread the needle. (The eye of a needle can refer to a parable that Jesus is reported to have told.)
- Any small gap can be called an eye.
- The middle of a hurricane is also called an eye – hence a calm part of a storm is called an eye. See eyewall.
- Eye teeth refers to teeth which are highest in the mouth.
- The dimples on the outside of a potato are referred to as eyes.
- Eye Weekly is an alternative newsweekly newspaper in Toronto, Canada
- Eyes (TV series) is the name of an American Broadcasting Company show that premired April 2005.
- "Eyes" is an episode of the science fiction television series Babylon 5; see Eyes (Babylon 5).
- Some herding dogs control herd animals using eye.
- "Eye" is a song by The Smashing Pumpkins featuring on the Lost Highway soundtrack.
- Eye of Horus - an Ancient Egyptian symbol of protection and power.
- Seeing Eye - guide dog school.
- Evil eye - widely distributed element of folklore or superstition.
- Eye of Providence - symbol showing an eye surrounded by rays of light or a glory, and usually enclosed by a triangle.
- In Sailing, the Cunningham's eye is one of the most important controls on a sail
- Eye is an abbreviation for the magazine-newspaper Private Eye
- The Eye (radio station), broadcasting from Melton Mowbray in England. Locations in the United Kingdom:
- Eye, Herefordshire, England
- Eye, Peterborough, England
- Eye, Suffolk, England
- Eye, Cambridgeshire, England
- Eye Green, Peterborough, England
- River Eye, England
- River Eye, Scotland Locations in the United States:
- Blue Eye, Arkansas
- Blue Eye, Missouri
- Red Eye Township, Minnesota
- Sleepy Eye, Minnesota Locations in Spain:
- Eyes (Andalucia)

Light

Light is electromagnetic radiation with a wavelength that is visible to the eye (visible light) or, in a technical or scientific context, electromagnetic radiation of any wavelength. The three basic dimensions of light (i.e., all electromagnetic radiation) are:
- Intensity (or brilliance or amplitude), which is related to the human perception of brightness of the light,
- Frequency (or wavelength), perceived by humans as the color of the light, and
- Polarization (or angle of vibration), which is not perceptible by humans under ordinary circumstances. Due to wave-particle duality, light simultaneously exhibits properties of both waves and particles. The precise nature of light is one of the key questions of modern physics.

Visible electromagnetic radiation

Visible light is the portion of the electromagnetic spectrum between the frequencies of 380 THz (3.8×10¹⁴ hertz) and 750 THz (7.5×10¹⁴ hertz). The speed (

c

), frequency (

f

\nu

), and wavelength (

\lambda

) of a wave obey the relation: :

c = f~\lambda \,\!

Because the speed of light in a vacuum is fixed, visible light can also be characterised by its wavelength of between 400 nanometres (abbreviated 'nm') and 800 nm (in a vacuum). Light entering the eye is absorbed by light-sensitive pigments within the rod cells and cone cells in the retina, triggering a cascade of events that creates electrical nerve impulses that travel through the optic nerve to the brain, producing vision.

Speed of light

Although some people speak of the "velocity of light", the word velocity should be reserved for vector quantities, that is, those with both magnitude and direction. The speed of light is a scalar quantity, having only magnitude and no direction, and therefore speed is the correct term. The speed of light has been measured many times, by many physicists. The best early measurement is Ole Rømer's (a Danish physicist), in 1676. By observing the motions of Jupiter and one of its moons, Io, with a telescope, and noting discrepancies in the apparent period of Io's orbit, Rømer calculated a speed of 227,000 kilometres per second (approximately 141,050 miles per second). The first successful measurement of the speed of light using an earthbound apparatus was carried out by Hippolyte Fizeau in 1849. Fizeau directed a beam of light at a mirror several thousand metres away, and placed a rotating cog wheel in the path of the beam from the source to the mirror and back again. At a certain rate of rotation, the beam could pass through one gap in the wheel on the way out and the next gap on the way back. Knowing the distance to the mirror, the number of teeth on the wheel, and the rate of rotation, Fizeau measured the speed of light as 313,000 kilometres per second. Léon Foucault used rotating mirrors to obtain a value of 298,000 km/s (about 185,000 miles/s) in 1862. Albert A. Michelson conducted experiments on the speed of light from 1877 until his death in 1931. He refined Foucault's results in 1926 using improved rotating mirrors to measure the time it took light to make a round trip from Mt. Wilson to Mt. San Antonio in California. The precise measurements yielded a speed of 186,285 mile/s (299,796 km/s [1,079,265,600 km/h]). In daily use, the figures are rounded off to 300,000 km/s and 186,000 miles/s.

Refraction

All light propagates at a finite speed. Even moving observers always measure the same value of c, the speed of light in vacuum, as c = 299,792,458 metres per second (186,282.397 miles per second). When light passes through a transparent substance, such as air, water or glass, its speed is reduced, and it undergoes refraction. The reduction of the speed of light in a denser material can be indicated by the refractive index, n, which is defined as: :

n = \frac \;\!

Thus, n=1 in a vacuum and n>1 in matter. When a beam of light enters a medium from vacuum or another medium, it keeps the same frequency and changes its wavelength. If the incident beam is not orthogonal to the edge between the media, the direction of the beam will change. Refraction of light by lenses is used to focus light in magnifying glasses, spectacles and contact lenses, microscopes and refracting telescopes.

Optics

The study of light and the interaction of light and matter is termed optics. The observation and study of optical phenomena such as rainbows offers many clues as to the nature of light as well as much enjoyment.

Color and wavelengths

The different wavelengths are detected by the human eye and then interpreted by the brain as colors, ranging from red at the longest wavelengths of about 700 nm. (lowest frequencies) to violet at the shortest wavelengths of about 400 nm. (highest frequencies). The intervening frequencies are seen as orange, yellow, green, cyan, blue, and, conventionally, indigo.
indigo
The wavelengths of the electromagnetic spectrum immediately outside the range that the human eye is able to perceive are called ultraviolet (UV) at the short wavelength (high frequency) end and infrared (IR) at the long wavelength (low frequency) end. Some animals, such as bees, can see UV radiation while others, such as pit viper snakes, can see infrared light. UV radiation is not normally directly perceived by humans except in a very delayed fashion, as overexposure of the skin to UV light can cause sunburn, or skin cancer, and underexposure can cause vitamin D deficiency. However, because UV is a higher frequency radiation than visible light, it very easily can cause materials to fluoresce visible light. Cameras that can detect IR and convert it to light are called, depending on their application, night-vision cameras or infrared cameras. These are different from image intensifier cameras, which only amplify available visible light. When intense radiation (of any frequency) is absorbed in the skin, it causes heating which can be felt. Since hot objects are strong sources of infrared radiation, IR radiation is commonly associated with this sensation. Any intense radiation that can be absorbed in the skin will have the same effect, however.

Measurement of light

The following quantities and units are used to measure the quantity or "brightness" of light. Light can also be characterised by:
- amplitude,
- color, wavelength, or frequency, and
- polarization (or angle of vibration).

Light sources

polarization There are many sources of light. The most common light sources are thermal: a body at a given temperature emits a characteristic spectrum of black body radiation. Examples include sunlight (the radiation emitted by the chromosphere of the Sun at around 6,000 K peaks in the visible region of the electromagnetic spectrum), incandescent light bulbs (which emit only around 10% of their energy as visible light and the remainder as infrared), and glowing solid particles in flames. The peak of the blackbody spectrum is in the infrared for relatively cool objects like human beings. As the temperature increases, the peak shifts to shorter wavelengths, producing first a red glow, then a white one, and finally a blue color as the peak moves out of the visible part of the spectrum and into the ultraviolet. These colors can be seen when metal is heated to "red hot" or "white hot". The blue color is most commonly seen in a gas flame or a welder's torch. Atoms emit and absorb light at characteristic energies. This produces "emission lines" in the spectrum of each atom. Emission can be spontaneous, as in light-emitting diodes, gas discharge lamps (such as neon lamps and neon signs, mercury-vapor lamps, etc.), and flames (light from the hot gas itself—so, for example, sodium in a gas flame emits characteristic yellow light). Emission can also be be stimulated, as in a laser or a microwave maser. Acceleration of a free charged particle, such as an electron, can produce visible radiation: cyclotron radiation, synchrotron radiation, and bremsstrahlung radiation are all examples of this. Particles moving through a medium faster than the speed of light in that medium can produce visible Cherenkov radiation. Certain chemicals produce visible radiation by chemoluminescence. In living things, this process is called bioluminescence. For example, fireflies produce light by this means, and boats moving through water can disturb plankton which produce a glowing wake. Certain substances produce light when they are illuminated by more energetic radiation, a process known as fluorescence. This is used in fluorescent lights. Some substances emit light slowly after excitation by more energetic radiation. This is known as phosphorescence. Phosphorescent materials can also be excited by bombarding them with subatomic particles. Cathodoluminescence is one example of this. This mechanism is used in cathode ray tube televisions. Certain other mechanisms can produce light:
- scintillation
- scintillator
- electroluminescence
- sonoluminescence
- triboluminescence
- radioactive decay
- particle-antiparticle annihilation

Theories about light

Early Greek ideas

In 55 BC Lucretius, continuing the ideas of earlier atomists, wrote that light and heat from the Sun were composed of minute particles. Ptolemy also wrote about the refraction of light.

10th century optical theory

The scientist Abu Ali al-Hasan ibn al-Haytham (965-c.1040), also known as Alhazen, developed a broad theory that explained vision, using geometry and anatomy, which stated that each point on an illuminated area or object radiates light rays in every direction, but that only one ray from each point, which strikes the eye perpendicularly, can be seen. The other rays strike at different angles and are not seen. He used the example of the pinhole camera, which produces an inverted image, to support his argument. Alhazen held light rays to be streams of minute particles that travelled at a finite speed. He improved Ptolemy's theory of the refraction of light. Alhazen's work did not become known in Europe until the late 16th century.

The 'plenum'

René Descartes (1596-1650) held that light was a disturbance of the plenum, the continuous substance of which the universe was composed. In 1637 he published a theory of the refraction of light which wrongly assumed that light travelled faster in a denser medium, by analogy with the behaviour of sound waves. Descartes' theory is often regarded as the forerunner of the wave theory of light.

Particle theory

Pierre Gassendi (1592-1655), an atomist, proposed a particle theory of light which was published posthumously in the 1660s. Isaac Newton studied Gassendi's work at an early age, and preferred his view to Descartes' theory of the plenum. He stated in his Hypothesis of Light of 1675 that light was composed of corpuscles (particles of matter) which were emitted in all directions from a source. One of Newton's arguments against the wave nature of light was that waves were known to bend around obstacles, while light travelled only in straight lines. He did, however, explain the phenomenon of the diffraction of light (which had been observed by Francesco Grimaldi) by allowing that a light particle could create a localised wave in the aether. Newton's theory could be used to predict the reflection of light, but could only explain refraction by incorrectly assuming that light accelerated upon entering a denser medium because the gravitational pull was greater. Newton published the final version of his theory in his Opticks of 1704. His reputation helped the particle theory of light to dominate physics during the 18th century.

Wave theory

In the 1660s, Robert Hooke published a wave theory of light. Christian Huygens worked out his own wave theory of light in 1678, and published it in his Treatise on light in 1690. He proposed that light was emitted in all directions as a series of waves in a medium called the aether. As waves are not affected by gravity, it was assumed that they slowed down upon entering a denser medium. The wave theory predicted that light waves could interfere with each other like sound waves (as noted in the 18th century by Thomas Young), and that light could be polarized. Young showed by means of a diffraction experiment that light behaved as waves. He also proposed that different colors were caused by different wavelengths of light, and explained color vision in terms of three-colored receptors in the eye. Another supporter of the wave theory was Euler. He argued in Nova theoria lucis et colorum (1746) that diffraction could more easily be explained by a wave theory. Later, Fresnel independently worked out his own wave theory of light, and presented it to the Académie des Sciences in 1817. Simeon Denis Poisson added to Fresnel's mathematical work to produce a convincing argument in favour of the wave theory, helping to overturn Newton's corpuscular theory. The weakness of the wave theory was that light waves, like sound waves, would need a medium for transmission. A hypothetical substance called the luminiferous aether was proposed, but its existence was cast into strong doubt by the Michelson-Morley experiment. Newton's corpuscular theory implied that light would travel faster in a denser medium, while the wave theory of Huygens and others implied the opposite. At that time, the speed of light could not be measured accurately enough to decide which theory was correct. The first to make a sufficiently accurate measurement was Léon Foucault, in 1850. His result supported the wave theory, and the classical particle theory was finally abandoned.

Electromagnetic theory

In 1845, Faraday discovered that the angle of polarisation of a beam of light as it passed through a polarising material could be altered by a magnetic field, an effect now known as Faraday rotation. This was the first evidence that light was related to electromagnetism. Faraday proposed in 1847 that light was a high-frequency electromagnetic vibration, which could propagate even in the absence of a medium such as the aether. Faraday's work inspired James Clerk Maxwell to study electromagnetic radiation and light. Maxwell discovered that self-propagating electromagnetic waves would travel through space at a constant speed, which happened to be equal to the previously measured speed of light. From this, Maxwell concluded that light was a form of electromagnetic radiation: he first stated this result in 1862 in On Physical Lines of Force. In 1873, he published A Treatise on Electricity and Magnetism, which contained a full mathematical description of the behaviour of electric and magnetic fields, still known as Maxwell's equations. The technology of radio transmission was, and still is, based on this theory. The constant speed of light predicted by Maxwell's equations contradicted the mechanical laws of motion that had been unchallenged since the time of Galileo, which stated that all speeds were relative to the speed of the observer. A solution to this contradiction would later be found by Albert Einstein.

Particle theory revisited

The wave theory was accepted until the late 19th century, when Einstein described the photoelectric effect, by which light striking a surface caused electrons to change their momentum, which indicated a particle-like nature of light. This clearly contradicted the wave theory, and for years physicists tried in vain to resolve this contradiction.

Quantum theory

In 1900, Max Planck described quantum theory, in which light is considered to be as a particle that could exist in discrete amounts of energy only. These packets were called quanta, and the particle of light was given the name photon, to correspond with other particles being described around this time, such as the electron and proton. A photon has an energy, E, proportional to its frequency, f, by :

E_f = hf = \frac \,\!

where h is Planck's constant,

\lambda

is the wavelength and c is the speed of light. As it originally stood, this theory did not explain the simultaneous wave-like nature of light, though Planck would later work on theories that did. The Nobel Committee awarded Planck the Physics Prize in 1918 for his part in the founding of quantum theory.

Wave-particle duality

The modern theory that explains the nature of light is wave-particle duality, described by Albert Einstein in the early 1900s, based on his work on the photoelectric effect and Planck's results. Einstein determined that the energy of a photon is proportional to its frequency. More generally, the theory states that everything has both a particle nature and a wave nature, and various experiments can be done to bring out one or the other. The particle nature is more easily discerned if an object has a large mass, so it took until an experiment by Louis de Broglie in 1924 to realise that electrons also exhibited wave-particle duality. Einstein received the Nobel Prize in 1921 for his work with the wave-particle duality on photons, and de Broglie followed in 1929 for his extension to other particles.

A light wave

1929 that oscillate perpendicular to each other and to the direction of motion (a transverse wave).]] The electric and magnetic fields are perpendicular to the direction of travel and to each other. This picture depicts a very special case, linearly polarized light. See Polarization for a description of the general case and an explanation of linear polarization. While these relations of the electric and magnetic fields are always true, the subtle difference in the general case is that the direction and amplitude of the magnetic (or electric) field can vary, in one place, with time, or, in one instant, can vary along the direction of propagation.

Sense

:This article is about the senses of living organsims (vision, taste, etc.). For other uses of the term, see sense (disambiguation). Senses are the physiological methods of perception. The senses and their operation, classification, and theory are overlapping topics studied by a variety of fields, most notably neuroscience, cognitive psychology (or cognitive science), and philosophy of perception.

Definition of "sense"

There is no firm agreement among neurologists as to exactly how many senses there are, because of differing definitions of a sense. In general, one can say that a "sense" is a faculty by which outside stimuli are perceived. School children are routinely taught that there are five senses (sight, hearing, touch, smell, taste; a classification first devised by Aristotle), it is generally agreed that there are at least nine different senses in humans, and a minimum of two more observed in other organisms. A broadly acceptable definition of a sense would be "a system that consists of a sensory cell type (or group of cell types) that respond to a specific kind of physical energy, and that correspond to a defined region (or group of regions) within the brain where the signals are received and interpreted." Where disputes arise is with regard to the exact classification of the various cell types and their mapping to regions of the brain.

List of Human senses

Using this definition several senses can be identified. Based on this outline and depending on the chosen method of classification, somewhere between 9 and 21 human senses have been identified. In addition, there are some other candidate physiological experiences which may or may not fall within the above classification (for example the sensory awareness of hunger and thirst).

Special senses

Sight or vision describes the ability to detect electromagnetic energy within the visible range (light) by the eye and the brain to interpret the image as "sight." There is disagreement as to whether this constitutes one, two or even three distinct senses. Neuroanatomists generally regard it as two senses, given that different receptors are responsible for the perception of colour (the frequency of photons of light) and brightness (amplitude/intensity - number of photons of light). Some argue that the perception of depth also constitutes a sense, but it is generally regarded that this is really a cognitive (that is, post-sensory) function of brain to interpret sensory input to derive new information. Hearing or audition is the sense of sound perception and results from tiny hair fibres in the inner ear detecting the motion of a membrane which vibrates in response to changes in the pressure exerted by atmospheric particles within (at best) a range of 9 to 20000 Hz, however this changes for each individual. Sound can also be detected as vibrations conducted through the body by tactition. Lower and higher frequencies than can be heard are detected this way only. Taste or gustation is one of the two main "chemical" senses. It is well-known that there are at least four types of taste "bud" (receptor) on the tongue and hence, as should now be expected, there are anatomists who argue that these in fact constitute four or more different senses, given that each receptor conveys information to a slightly different region of the brain. The four well-known receptors detect sweet, salt, sour, and bitter, although the receptors for sweet and bitter have not been conclusively identified. A fifth receptor, for a sensation called umami, was first theorised in 1908 and its existence confirmed in 2000 (see [http://www.nature.com/neuro/press_release/nn0200.html]). The umami receptor detects the amino acid glutamate, a flavor commonly found in meat, and in artificial flavourings such as monosodium glutamate. Smell or olfaction is the other "chemical" sense. Unlike taste, there are hundreds of olfactory receptors, each binding to a particular molecular feature, according to current theory. The combination of features of the odor molecule makes up what we perceive as the molecule's smell. In the brain, olfaction is processed by the olfactory system. Olfactory receptor neurons in the nose differ from most other neurons in that they die and regenerate on a regular basis. If the different taste-senses are not regarded as separate senses one may argue that Taste and Smell should likewise be grouped together as one sense.

Somatic senses

Touch or tactition is the sense of pressure perception, generally in the skin. There are a variety of pressure receptors that respond to variations in pressure (firm, brushing, sustained, etc). Thermoception is the sense of heat and the absence of heat (cold), also by the skin and including internal skin passages. There is some disagreement about how many senses this actually represents - the thermoceptors in the skin are quite different from the homeostatic thermoceptors which provide feedback on internal body temperature. Nociception is the perception of pain. It can be classified as from one to three senses, depending on the classification method. The three types of pain receptors are cutaneous (skin), somatic (joints and bones) and visceral (body organs). For a considerable time, it was believed that pain was simply the overloading of pressure receptors, but research in the first half of the 20th century indicated that pain is a distinct phenomenon that intertwines with all other senses, including touch.

Other

Equilibrioception is the perception of balance and is related to cavities containing fluid in the inner ear. There is some disagreement as to whether this also includes the sense of "direction" or orientation. However, as with depth perception earlier, it is generally regarded that "direction" is a post-sensory cognitive awareness. Proprioception is the perception of body awareness and is a sense that people rely on enormously, yet are frequently not aware of. More easily demonstrated than explained, proprioception is the "unconscious" awareness of where the various regions of the body are located at any one time. (This can be demonstrated by anyone's closing the eyes and waving the hand around. Assuming proper proprioceptive function, at no time will the person lose awareness of where the hand actually is, even though it is not being detected by any of the other senses).

Non-human senses

Other living organisms have receptors to sense the world around them, including many of the senses listed above for humans. However, the mechanisms and capabilities vary widely. Among non-human animals, dogs have a much keener sense of smell than humans, although the mechanism is similar. Pit vipers and some boas have organs that allow them to detect infrared light, such that these snakes are able to sense the body heat of their prey. This is, however, also just sight extended to include more frequencies. Insects have olfactory receptors on their antennae. Ctenophores have a balance receptor (a statocyst) that works very differently from the mammalian semi-circular canals. In addition, some animals have senses that humans do not, including the following: Electroception (or "electroreception"), the most significant of the non-human senses, is the ability to detect electric fields. Several species of fish, sharks and rays have evolved the capacity to sense changes in electric fields in their immediate vicinity. Some fish passively sense changing nearby electric fields, some generate their own weak, electric fields and sense the pattern of field potentials over their body surface, and some use these generating and sensing capacities for social communication. The mechanisms by which electroceptive fishes construct a spatial representation from very small differences in field potentials involve comparisons of spike latencies from different parts of the fish's body. The only order of mammals that is known to demonstrate electroception is the monotreme order. Among these mammals, the platypus (see [http://web.archive.org/web/19981206164009/http://instruct1.cit.cornell.edu/courses/bionb420.07/anelson/platypus.html]) has the most acute sense of electroception. Humans (and probably other mammals) can detect electric fields indirectly by detecting the effect they have on hairs. An electrically charged balloon, for instance, will exert a force on human arm hairs, which can be felt through tactition and identified as coming from a static charge (and not from wind or the like). This is however not Electroception since there is no separate sense for it. The presence of an electrical field is merely concluded from a side-effect of another sense. Magnetoception (or "magnetoreception") is the ability to detect fluctuations in magnetic fields and is most commonly observed in birds, though it has also been observed in insects such as bees. Although there is no dispute that this sense exists in many avians (it is essential to the navigational abilities of migratory birds), it is not a well-understood phenomenon (see [http://www.ks.uiuc.edu/Research/magsense/ms.html]). Magnetotactic bacteria build miniature magnets inside themselves and use them to determine their orientation relative to the Earth's magnetic field. Echolocation is the ability to determine orientation to other objects through interpretation of reflected sound (like sonar). Bats and cetaceans are noted for this ability, though some other animals use it, as well. It is most often used to navigate through poor lighting conditions or to identify and track prey. There is presently an uncertainty whether this is simply an extremely developed post-sensory interpretation of auditory perceptions or it actually constitutes a separate sense. Resolution of the issue will require brain scans of animals while they actually perform echolocation, a task that has proven difficult in practice. Pressure detection uses the lateral line, which is a pressure-sensing system of hairs found in fish and some aquatic amphibians. It is used primary for navigation, hunting, and schooling. In a way, plants and some microorganisms can sense, too. For example, the mimosa plant is popular for folding up its leaves as soon as one touches it.

External links

- [http://samvak.tripod.com/sense.html The physiological and psychological underpinnings of senses]
- [http://www.med.uwo.ca/physiology/courses/sensesweb The Physiology of the Senses tutorial] 12 animated chapters on vision, hearing, touch, balance and memory.
- The 2004 Nobel Prize in Physiology or Medicine ([http://nobelprize.org/medicine/laureates/2004/index.html announced] 4 October 2004) was won by Richard Axel and Linda Buck for their work explaining olfaction, published first in a joint paper in 1991 that described the very large family of about one thousand genes for odorant receptors and how the receptors link to the brain. Category:Sensory system ja:五感 simple:Sense

Visual perception

Visual perception is one of the senses, consisting of the ability to detect light and interpret (see) it as the perception known as sight or naked eye vision. Vision has a specific sensory system, the visual system. There is disagreement as to whether or not this constitutes one, two or even three distinct senses. Some people make a distinction between "black and white" vision and the perception of colour, and others point out that vision using rod cells uses different physical detectors on the retina from cone cells. Some argue that the perception of depth also constitutes a sense, but others argue that this is really cognition (that is, post-sensory) function derived from having stereoscopic vision (two eyes) and is not a sensory perception as such. Many people are also able to perceive the polarization of light.

The visual system

thumbnailThe eye is the light-sensitive organ that is the first component of the visual system. The eye's retina performs the first stages of visual perception processing, with the remaining stages of visual perception occurring in the optic nerve, the lateral geniculate nucleus, and the visual cortex of the brain.

Sources of information

To perform its task, visual perception takes into account not only patterns of illumination on the retina, but also our other senses and our past experiences. Consider the task of bird sighting (an instance of object recognition): to be able to identify a bird against a background of tree and brushes, one needs prior exposure to general properties of the bird category. From past experiences, we expect birds to have a certain shape, color, etc. Hearing a sound that is characteristic of birds, a song for example, will help us locate one: information from the other senses is used in visual perception. In this case, locational information from the auditory domain is used.

Individual and group differences in visual perception

Most of the general processes of visual perception have been shown to be universal, as opposed to being dependant on culture, although there are specific instances where cultural variability appears to come into play. It has also been shown that certain individual differences such as impairment of sight and spatial skills can also affect our visual perception. There are also other factors that influence how we perceive things such as personality, cognitive styles, gender, occupation, age, values, attitudes, motivation, religious beliefs, economic status, education and habits.

Theoretical perspectives in the study of visual perception

Unconscious inference

Hermann von Helmholtz is often credited with the founding of the scientific study of visual perception. Helmholtz held vision to be a form of unconscious inference: vision is a matter of deriving a probable interpretation for incomplete data. The general goal of vision is to identify, as accurately as possible, the features of our environment: roughly, what objects are present where. Other features are irrelevant to this task : illumination patterns, viewing position, etc. Those are confounding variables. Call S = (F,C) the scene, with F the features we’re interested in and C the confounding variables. S determines I, the pattern of illumination on the retina, which is all the information our visual system has on the current scene. The task is to find S given I. This problem is under-constrained: many different S correspond to the same I, and many I could correspond to the same S. One of the reasons is that much information is lost when a 3-dimensional world is collapsed into a 2-dimensional array. To see why, consider the figure of a circle such as this one: O. It could correspond to an infinity of ellipses viewed at a certain slant. But we always interpret it as a circle viewed on the frontal plane – the explanation we infer from the data for this particular stimulus. Inference requires prior assumptions about the world: two well-known assumptions that we make in processing visual information are that light comes from above and that objects are viewed from above not below. The study of visual illusions (cases when the inference process goes wrong) has yielded a lot of insight into what sort of assumptions the visual system makes.

Gestalt

Psychologists of the Gestalt school have raised a large part of the research questions that still preoccupy vision scientists today. The so-called Gestalt Laws of Organisation have broadened the study of how people perceive objects to be organized patterns or wholes, instead of collections of many separate parts. Gestalt is a German word that translates to "configuration or pattern". According to this theory, there are four main factors that determine how we group things according to visual perception.
- Proximity – Depending on how close object are to one other, we tend to group the ones closest to each other as a group.
- Similarity – If objects are similar in shape or size to one another we tend to group them together.
- Closure – How we complete a pattern because of how the items are grouped together even though the pattern is not complete.
- Simplicity – How we group items according to symmetry, regularity, and smoothness.

Ecological psychology

Psychologist James J. Gibson developed a theoretical perspective on vision that is radically different from that of Helmholtz. Gibson considers that enough visual perception is available in normal environments to allow for veridical perception (accurate perception of the world). Gibson replaces inference with information pickup. Although most researchers today feel closer to Helmholtz's unconscious inference theory, Gibson has done much in identifying what sort of information is available to the visual system.

Types of visual perception

- Black and white vision
- Color vision
- Gestalt perception
- Motion perception

Disorders/Dysfuntions

- Achromatopsia
- Color blindness
- Scotopic Sensitivity Syndrome

References

- Rudolph Arnheim (1954). Art and Visual Perception: A Psychology of the Creative Eye. Berkeley: University of California Press.
- Lothar Kleine-Horst (2001). Empiristic Theory of Visual Gestalt Perception. Hierarchy and Interactions of Visual Functions. Koeln: Enane. ISBN 3-928955-42X

External links

- [http://enane.de/cont.htm Empiristic theory of visual gestalt perception]
- [http://www.aber.ac.uk/media/Modules/MC10220/visper03.html Visual Perception 3 - Cultural and Environmental Factors]
- [http://www.sapdesignguild.org/resources/optical_illusions/gestalt_laws.html Gestalt Laws]
- [http://www.aber.ac.uk/media/Modules/MC10220/visper04.html Visual Perception 4 - Individual Differences, Purposes and Needs] Category:Computer vision Category:Vision ja:視覚

Organism

In biology and ecology, an organism (in Greek organon = instrument) is a complex adaptive system of organs that influence each other in such a way that they function as a more or less stable whole and have properties of life. The origin of life and the relationships between its major lineages are controversial. Two main grades may be distinguished, the prokaryotes and eukaryotes. The prokaryotes are generally considered to represent two separate domains, called the Bacteria and Archaea, which are not closer to one another than to the eukaryotes. The gap between prokaryotes and eukaryotes is widely considered a major missing link in evolutionary history. Two eukaryotic organelles, namely mitochondria and chloroplasts, are generally considered to be derived from endosymbiotic bacteria. The phrase complex organism describes any organism with more than one cell.

Organizational terminology

Biological Organization

Viruses

Viruses are not typically considered to be organisms because they are not capable of independent reproduction or metabolism. However, according to the United States Code, they are considered to be microorganisms in the sense of biological weaponry and malicious use. This controversy is problematic, though, since some parasites and endosymbionts are incapable of independent life either. Although viruses do have enzymes and molecules characteristic of living organisms, they are incapable of surviving outside a host cell and most of their metabolic processes require a host and its 'genetic machinery'. The origin of such parasites is uncertain, but it appears most likely that they are derived from their host.

Life span

One of the basic parameters of organism is its life span. Some animals live as short as one day, while some plants can live thousands of years. Aging is important when determining life span of most organisms, bacterium, a virus or even a prion.

External links

- [http://news.bbc.co.uk/1/hi/sci/tech/944790.stm BBCNews: 27 September, 2000, When slime is not so thick] Citat: "...It means that some of the lowliest creatures in the plant and animal kingdoms, such as slime and amoeba, may not be as primitive as once thought...."
- [http://www.spaceref.com/news/viewpr.html?pid=4742 SpaceRef.com, July 29, 1997: Scientists Discover Methane Ice Worms On Gulf Of Mexico Sea Floor]
- [http://www.science.psu.edu/iceworms/iceworms.html The Eberly College of Science: Methane Ice Worms discovered on Gulf of Mexico Sea Floor] download Publication quality photos
- [http://www.sb-roscoff.fr/Ecophy/PDF/00-Fisher-NatWis.pdf Artikel, 2000: Methane Ice Worms: Hesiocaeca methanicola. Colonizing Fossil Fuel Reserves]
- [http://www.spaceref.com/news/viewnews.html?id=339 SpaceRef.com, May 04, 2001: Redefining "Life as We Know it"] Hesiocaeca methanicola In 1997, Charles Fisher, professor of biology at Penn State, discovered this remarkable creature living on mounds of methane ice under half a mile of ocean on the floor of the Gulf of Mexico.
- [http://news.bbc.co.uk/1/hi/sci/tech/2585235.stm BBCNews, 18 December, 2002, 'Space bugs' grown in lab] Citat: "...Bacillus simplex and Staphylococcus pasteuri...Engyodontium album...The strains cultured by Dr Wainwright seemed to be resistant to the effects of UV - one quality required for survival in space...."
- [http://news.bbc.co.uk/1/hi/sci/tech/3003946.stm BBCNews, 19 June, 2003, Ancient organism challenges cell evolution] Citat: "..."It appears that this organelle has been conserved in evolution from prokaryotes to eukaryotes, since it is present in both,"..."
- [http://www.anselm.edu/homepage/jpitocch/genbios/bi04syllabsu03.html Interactive Syllabus for General Biology - BI 04, Saint Anselm College, Summer 2003]
- [http://www.personal.psu.edu/users/j/s/jsf165/Bio110.html Jacob Feldman: Stramenopila]
- [http://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Root NCBI Taxonomy entry: root] (rich)
- [http://www.anselm.edu/homepage/jpitocch/genbios/surveybi04.html Saint Anselm College: Survey of representatives of the major Kingdoms] Citat: "...Number of kingdoms has not been resolved...Bacteria present a problem with their diversity...Protista present a problem with their diversity...",
- [http://www.species2000.org/ Species 2000 Indexing the world's known species]. Species 2000 has the objective of enumerating all known species of plants, animals, fungi and microbes on Earth as the baseline dataset for studies of global biodiversity. It will also provide a simple access point enabling users to link from here to other data systems for all groups of organisms, using direct species-links.
- [http://www.abc.net.au/science/news/enviro/EnviroRepublish_828525.htm The largest organism in the world may be a fungus carpeting nearly 10 square kilometers of an Oregon forest, and may be as old as 8500 years.]
- [http://tolweb.org/tree/phylogeny.html The Tree of Life]. zh-min-nan:Seng-bu̍t ko:생물 ja:生物 th:สิ่งมีชีวิต

Bird

Many - see section below. Birds are bipedal, warm-blooded, egg-laying vertebrates characterized primarily by feathers, forelimbs modified as wings, and hollow bones. Birds range in size from the tiny hummingbirds to the huge Ostrich and Emu. Depending on taxonomic viewpoint, there are about 8,800–10,200 living bird species (plus about 120–130 that have become extinct in the span of human history) in the world, making them the most diverse class of terrestrial vertebrates. Birds are a very differentiated class, with some feeding on nectar, plants, seeds, insects, rodents, fish, carrion, or other birds. Most birds are diurnal, or active during the day. Some birds, such as the owls and nightjars, are nocturnal or crepuscular (active during twilight hours). Many birds migrate long distances to utilise optimum habitats (e.g., Arctic Tern) while others spend almost all their time at sea (e.g. the Wandering Albatross). Some, such as frigatebirds, stay aloft for days at a time, even sleeping on the wing. Common characteristics of birds include a bony beak with no teeth, the laying of hard-shelled eggs, high metabolic rate, and a light but strong skeletons. Most birds are characterised by flight, though the ratites are flightless, and several other species, particularly on islands, have also lost this ability. Flightless birds include the penguins, Ostrich, kiwi, and the extinct Dodo. Flightless species are vulnerable to extinction when humans or the mammals they introduce arrive in their habitat, for example the Great Auk, flightless rails, and the moa of New Zealand.

Bird orders

New Zealand This is a list of the taxonomic orders in the class Aves. The list of birds gives a more detailed summary, including families.
- Struthioniformes, Ostrich, emus, kiwis, and allies
- Tinamiformes, tinamous
- Anseriformes, waterfowl
- Galliformes, fowl
- Sphenisciformes, penguins
- Gaviiformes, loons
- Podicipediformes, grebes
- Procellariiformes, albatrosses, petrels, and allies
- Pelecaniformes, pelicans and allies
- Ciconiiformes, storks and allies
- Phoenicopteriformes, flamingos
- Accipitriformes, eagles, hawks and allies
- Falconiformes, falcons
- Turniciformes, button-quail
- Gruiformes, cranes and allies
- Charadriiformes, plovers and allies
- Pteroclidiformes, sandgrouse
- Columbiformes, doves and pigeons
- Psittaciformes, parrots and allies
- Cuculiformes, cuckoos
- Strigiformes, owls
- Caprimulgiformes, nightjars and allies
- Apodiformes, swifts
- Trochiliformes, hummingbirds
- Coraciiformes, kingfishers
- Piciformes, woodpeckers and allies
- Trogoniformes, trogons
- Coliiformes, mousebirds
- Passeriformes, passerines Note: This is the traditional classification (the so-called Clements order). A more recent, radically different classification based on molecular data has been developed (the so-called Sibley order) and is gaining acceptance.

Evolution

Birds are generally considered to have evolved from theropod dinosaurs. Specifically, birds are members of Maniraptora, a group of theropods which includes dromaeosaurs and oviraptorids. As more non-avian theropods that are closely related to birds are discovered, the formerly clear distinction between non-birds and birds becomes less so. Recent discoveries in North-east China (Liaoning Province) demonstrating that many small theropod dinosaurs had feathers contribute to this ambiguity. The basal bird Archaeopteryx, from the Jurassic, is well-known as one of the first "missing links" to be found in support of evolution in the late 19th century. It remains the most primitive known bird. Other Mesozoic birds include the Confuciusornithidae, Enantiornithes, Ichthyornis, and Hesperornithiformes, a group of flightless divers resembling grebes and loons. The recently discovered dromaeosaur, Cryptovolans, was capable of powered flight, contained a keel and had ribs with uncinate processes. In fact, Cryptovolans makes a better "bird" than Archaeopteryx which is missing some of these modern bird features. Because of this, some paleontologists have suggested that dromaeosaurs are actually basal birds whose larger members are secondarily flightless, i.e. dromaeosaurs evolved from birds and not the other way around. Evidence for this theory is currently inconclusive, but digs continue to unearth fossils (especially in China) of the strange feathered dromaeosaurs. It should be noted that although ornithischian (bird-hipped) dinosaurs share the same hip structure as birds, birds actually originated from the saurischian (lizard-hipped) dinosaurs, and thus arrived at their hip structure condition independently. In fact, the bird-like hip structure developed a third time among a peculiar group of theropods, the Therizinosauridae. Modern birds are classified in Neornithes, which are split into the Paleognathae and Neognathae. The paleognaths include the tinamous (found only in Central and South America) and the ratites. The ratites are large flightless birds, and include ostriches, cassowaries, kiwis and emus; some scientists suspect that the ratites represent an artificial grouping of birds which have independently lost the ability to fly, others contend that the ratites never had the ability to fly and are more directly related to the dinosaurs. The basal divergence from the remaining Neognathes was that of the Galloanseri, the superorder containing the Anseriformes (ducks, geese and swans), and the Galliformes (the pheasants, grouse, and their allies). See the chart. The classification of birds is a contentious issue. Sibley & Ahlquist's Phylogeny and Classification of Birds (1990) is a landmark work on the classification of birds (although frequently debated and constantly revised). Evidence for the various orders seems to be fairly good, but the relationships between the orders are in a state of disarray. Evidence from modern bird anatomy, fossils and DNA have all been brought to bear on the problem but no strong consensus has emerged. See also: Sibley-Ahlquist taxonomy.

Reproduction

Although most male birds have no external sex organs, the male does have two testes which become hundreds of times larger during the breeding season to produce sperm. The female's ovaries also become larger, although only the left ovary actually functions. In the males of species without a phallus (see below), sperm is stored within the proctodeum compartment within the cloaca prior to copulation. During copulation, the female moves her tail to the side and the male either mounts the female from behind or moves very close to her. He moves the opening of his cloaca, or vent, close to hers, so that the sperm can enter the female's cloaca, in what is referred to as a cloacal kiss. This can happen very fast, sometimes in less than one second. The sperm is stored in the female's cloaca for anywhere from a week to a year, depending on the species of bird. Then, one by one, eggs will descend from the female's ovaries and become fertilized by the male's sperm, before being subsequently laid by the female. The eggs will then continue their development in the nest. cloacal kiss.]] Many waterfowl and some other birds, such as the ostrich and turkey, do possess a phallus. Except during copulation, it is hidden within the proctodeum compartment within the cloaca, just inside the vent. The avian phallus differs from the mammalian penis in several ways, most importantly in that it is purely a copulatory organ and is not used for dispelling urine. After the eggs hatch, parent birds provide varying degrees of care in terms of food and protection. Precocial birds can care for themselves independently within minutes of hatching; altricial hatchlings are helpless, blind, and naked, and require extended parental care. The chicks of many ground-nesting birds such as partridges and waders are often able to run virtually immediately after hatching; such birds are referred to as nidifugous. The young of hole-nesters, on the other hand, are often totally incapable of unassisted survival. "Fledging" is the process of a chick acquiring feathers until it can fly. Some birds, such as pigeons, geese, and Red-crowned Cranes, remain with their mates for life (or for a long period) and may produce offspring on a regular basis.

Mating systems and parental care

Sources for this section include:
- Gowaty, Patricia Adair: Male Parental Care and Apparent Monogamy among Eastern Bluebirds (Sialia Sialis). The American Naturalist 121(2): 149-160 (1983).
- Ketterson, Ellen D. and Nolan, Val: Male Parental Behavior in Birds. Annual Review of Ecology and Systematics 25: 601-28 (1994).
- Zeveloff, Samuel and Boyce, Mark: Parental Investment and Mating Systems in Mammals. Evolution 34(5): 973-982 (1980).

The three predominant mating systems are polyandry, polygyny, and monogamy. Monogamy is seen in approximately 91% of all bird species. Polygyny constitutes 2% of all birds and polyandry is seen in less than 1%. Monogamous species of males and females pair for the breeding season. In some cases, the individuals may pair for life. One reason for the high rate of monogamy among birds is due to the fact that male birds are just as adept at parental care as females. In most groups of animals, male parental care is rare, but in birds it is quite common; it is more extensive in birds than in any other vertebrate class. In fact, male care can be seen as important or essential to female fitness. "In one form of monogamy such as with obligate monogamy a female cannot rear a litter without the aid of a male" (Gowaty, 1983). obligate The parental behavior most associated with monogamy was male incubation. This is very interesting, because male incubation is the most confining male parental behavior. It not only consumes time, but also may require physiological changes that interfere with usual mating. With the extreme loss of mating