:: wikimiki.org ::

Kudzu Vine

Kudzu Vine

Kudzu, Pueraria lobata (syn. P. montana, P. thunbergiana), is one of about 20 species in the genus Pueraria in the pea family Fabaceae, subfamily Faboideae. It is native to southern Japan and southeast China in eastern Asia. The name comes from Japanese kuzu (葛), meaning vine. The other species of Pueraria also occur in southeast Asia, mostly further south. It is a climbing, woody or semi-woody, perennial vine capable of reaching heights of 20–30 m (66-98 ft) in trees, but also scrambles extensively over lower vegetation. The leaves are deciduous, alternate and compound, with a petiole (leaf stem) 10–20 cm (4–8 in) long and three broad leaflets 14–18 cm (6–7 in) long and 10 cm (4 in) broad. The leaflets may be entire or deeply 2–3 lobed, and are pubescent underneath with hairy margins. The flowers are borne in long panicles 10–25 cm (about 4–10 in) long with about 30–80 individual blooms at nodes on the stems (see image). Each flower is about 1–1.5 cm (about 0.4–0.6 in) long, purple, highly fragrant. The flowers are copious nectar producers and are visited by many species of insects, including bees, butterflies and moths. Flowering occurs in late summer and is soon followed by production of brown, hairy, flattened, seed pods, each of which contains three to ten hard seeds. The non-woody parts of the plant are edible. The young leaves can be used for salad or cooked as a leaf vegetable; the flowers battered and fried (like squash flowers); and the starchy tuberous roots can be prepared as any root vegetable. Once established, kudzu plants grow rapidly, extending as much as 20 m (60 ft) per season at a rate of about 30 cm (12 in) per day. This vigorous vine may extend 10–30 m (30–100 ft) in length, with basal stems 1–10 cm (1–4 in) diameter. Kudzu roots are fleshy, with massive tap roots 10–20 cm (4–8 in) or more in diameter, 1–2 m (3–6 ft) or more in length, and weighing as much as 180 kg. As many as thirty stems may grow from a single root crown. Kudzu grows well under a wide range of conditions and in most soil types. Preferred habitats are forest edges, abandoned fields, roadsides, and disturbed areas, where sunlight is abundant. Kudzu grows best where winters do not drop below –15 °C (5 °F), average summer temperatures are regularly above 27 °C (80 °F), and annual rainfall is 1000 mm (40 in) or more. In areas where winters drop below –15 °C, it will be killed to ground level, but the roots may send up new growth in the spring. Kudzu is sometimes referred to as "the plant that ate the South", a reference to how kudzu's explosive growth has been most prolific in the southeastern United States due to nearly ideal growing conditions. Significant sums of money and effort are spent each growing season to prevent kudzu from taking over roads, bridges, power lines, and local vegetation.

Uses

Culinary

The starchy roots are ground into a fine powder and used for varieties of Wagashi and herbal medicines. When added to water and heated, kudzu powder becomes clear and adds stickiness to the food. The name Kudzu appeared first in Kojiki and Nihonshoki as a type of vine or Kazura (葛 or 蔓) used commonly by the people who lived in Kudzu (国栖), area around present-day Yoshino, Nara prefecture. It is unclear whether the name was taken from the people or the name of the plant was applied to the people. Kudzu has been in use for over 1300 years and it is speculated that it goes back even further. In the Nara and Heian era, records had been found that they were collected and sent as a part of tax. Even today, "Yoshino Kudzu" has the best image of kudzu powder yet. Kagoshima prefecture is the largest producer of kudzu products.

Medicinal

Studies have shown ([http://www.med.unc.edu/alcohol/research/Overstreet/kudzu.html], [http://www.alcoholism-cer.com/pt/re/alcoholism/abstract.00000374-200505000-00008.htm;jsessionid=CiQY3kwMmK6wIZ8V9SUTyFS9L7C2ZbYFuVrJVOnd7601kcXkVYYa!-2006515172!-949856031!9001!-1?index=1&results=1&count=10&searchid=1&nav=search&database=lwwovft]) that kudzu can reduce both hangovers and alcohol cravings. The mechanism for this is not yet established, but it may have to do with both alcohol metabolism and the reward circuits in the brain. Kudzu also contains a number of useful isoflavones, including daidzein (an anti-inflammatory and antimicrobial agent), daidzin (a cancer preventive) and genistein (an antileukemic agent). Kudzu is a unique source of fourth isoflavone, Puerarin. Kudzu root compounds can affect neurotransmitters (including serotonin, GABA, and glutamate) and it has shown value in treating migraine and cluster headache [http://med-owl.com/clusterheadaches/tiki-index.php?page=Kudzu]. In traditional Chinese medicine, kudzu was used for tinnitus, vertigo, and Wei syndrome (superficial heat close to the surface).

Kudzu as an invasive exotic

Kudzu was introduced from Japan into the United States in 1876 at the Philadelphia Centennial Exposition, where it was promoted as a forage crop and an ornamental plant. From 1935 to the early 1950s, farmers in the South were encouraged to plant kudzu to reduce soil erosion, and Franklin D. Roosevelt's Civilian Conservation Corps planted it widely for many years. Kudzu was recognized as a pest weed by the United States Department of Agriculture in 1953, and was removed from its list of permissible cover plants. Kudzu is now common throughout most of the southeastern United States, and has been found as far north as Pennsylvania, and as far south as northern Florida. It has also been found growing (rather inexplicably) in Clackamas County, Oregon in 2000 ([http://www.oda.state.or.us/information/news/2000/kudzu.html]). In all, kudzu infests 20,000 to 30,000 square kilometres of land in the United States and costs around $500 million annually in lost cropland and control costs. It cannot tolerate extremely low freezing temperatures that bring the frost line down through its entire root system; however it does require some cold weather (a solid frost or freeze annually). Kudzu is also becoming a problem in northeastern Australia. Australia Kudzu vines can make walking across an area nearly impossible, as it takes over all horizontal and vertical surfaces, both natural and manmade. Its dense growth obstructs all views and movement into the area. It kills or degrades other plants by smothering them under a solid blanket of leaves, by girdling woody stems and tree trunks, and by breaking branches or uprooting entire trees and shrubs through its weight. The spread of kudzu is mainly by vegetative expansion by runners and rhizomes and by vines that root at the nodes to form new plants. Kudzu will also spread by seeds, which are contained in pods and mature in the autumn. One or two viable seeds are produced per cluster of pods. These hard-coated seeds may not germinate for several years, which can result in the re-appearance of the species years after it was thought eradicated at a site.

Control

For successful long-term control of kudzu, the extensive root system must be destroyed. Any remaining root crowns will lead to reinfestation of an area. Mechanical methods involve cutting vines just below ground level and destroying all cut material. Close mowing every month or regular heavy grazing for two growing seasons, or repeated cultivation may be effective. Cut kudzu can be fed to livestock, burned or enclosed in plastic bags and sent to a landfill. If done in the spring, cutting must be repeated as regrowth appears to exhaust the plant's stored carbohydrate reserves. Late-season cutting should be followed up with immediate application of a systemic herbicide to cut stems, to encourage transport of the herbicide into the root system. Repeated applications of several soil-active herbicides have been used effectively on large infestations in forestry situations. Efforts are being organized by the U.S. Forest Service to search for biological control agents for kudzu, and a particular fungus is currently in testing.

External links

- [http://www.cptr.ua.edu/kudzu/ The Amazing Story of Kudzu]
- [http://www.jjanthony.com/kudzu/ Pictures of Kudzu "consuming" cars and houses]
- [http://www.uptowncity.com/Dixie/khumor.htm How to grow Kudzu]
- [http://www.nps.gov/plants/alien/fact/pulo1.htm More technical details]
- [http://www.invasivespecies.gov/profiles/kudzu.shtml Invasivespecies.gov article] (lots of links)
- [http://www.kudzukingdom.com/ Uses for Kudzu] (a "pro-Kudzu" site)
- [http://www.scienceblog.com/cms/node/7921 Kudzu as anti-alcohol treatment]
- [http://www.med-owl.com/clusterheadaches/tiki-index.php?page=Kudzu Monograph on Medicinal Uses of Kudzu]
- [http://www2.potsdam.edu/hansondj/HealthIssues/1127332920.html Kudzu and Alcohol Consumption] Part of this article was originally based on content from public domain web pages from the United States National Park Service and the United States Bureau of Land Management. Category:Faboideae Category:Root vegetables Category:Invasive species Category:Edible thickening agents ja:クズ

Species

In biology, a species is the basic unit of biodiversity. In scientific classification, a species is assigned a two-part name in Latin. The genus is listed first (and capitalized), followed by a specific epithet. For example, humans belong to the genus Homo, and are in the species Homo sapiens. The name of the species is the whole binomial not just the second term (the specific epithet). The binomial, and most other purely formal aspects of the biological codes of nomenclature, were formalized by Carolus Linnaeus in the 1700's and as a result are called the "Linnaean system". At that time, species were thought to represent independent acts of creation by God, and were therefore considered objectively real and immutable. Since the advent of the theory of evolution, the conception of species has undergone vast changes in biology, however no consensus on the definition of the word has yet been reached. The most commonly cited definition of "species" was first coined by Ernst Mayr. By this definition, called the biological species concept or isolation species concept, species are "groups of actually or potentially interbreeding natural populations which are reproductively isolated from other such groups". However, many other species concepts are also used (see other definitions of species below). The scientific name of a species is properly typeset in italics. When an unknown species is being referred to this may be done by using the abbreviation "sp." in the singular or "spp." in the plural in the place of the second part of the scientific name. Note that the word "specie" is not the singular of "species". It refers to coined money.

Definitions of species

The definition of a species given above as taken from Mayr, is somewhat idealistic. Since it assumes sexual reproduction, it leaves the term undefined for a large class of organisms that reproduce asexually. Biologists frequently do not know whether two morphologically similar groups of organisms are "potentially" capable of interbreeding. Further, there is considerable variation in the degree to which hybridization may succeed under natural and experimental conditions, or even in the degree to which some organisms use sexual reproduction between individuals to breed. Consequently, several lines of thought in the definition of species exist: ; Typological species : A group of organisms in which individuals are members of the species if they sufficiently conform to certain fixed properties. The clusters of variations or phenotypes within specimens (ie: longer and shorter tails) would differentiate the species. This method was used as a "classical" method of determining species, such as with Linnaeus early in evolutionary theory. However, we now know that different phenotypes do not always constitute different species (e.g.: a 4-winged Drosophila born to a 2-winged mother is not a different species). Species named in this manner are called morphospecies. ; Morphological species : A population or group of populations that differs morphologically from other populations. For example, we can distinguish between a chicken and a duck because they have different shaped bills and the duck has webbed feet. Species have been defined in this way since well before the beginning of recorded history. This species concept is much criticised because more recent genetic data reveals that genetically distinct populations may look very similar and, contrarily, large morphological differences sometimes exist between very closely-related populations. Nonetheless, most species known have been described solely from morphology. ; Biological / Isolation species : A set of actually or potentially interbreeding populations. This is generally the most useful formulation for scientists working with living examples of the higher taxa like mammals, fish, and birds, but meaningless for organisms that do not reproduce sexually. It does not distinguish between the theoretical possibility of interbreeding and the actual likelihood of gene flow between populations and is thus impractical in instances of allopatric (geographically isolated) populations. The results of breeding experiments done in artificial conditions may or may not reflect what would happen if the same organisms encountered each other in the wild, making it difficult to gauge whether or not the results of such experiments are meaningful in reference to natural populations. ; Mate-recognition species : A group of organisms that are known to recognise one another as potential mates. Like the isolation species concept above, it applies only to organisms that reproduce sexually. Unlike the isolation species concept, it focuses specifically on pre-mating reproductive isolation. ; Phylogenetic / Evolutionary / Darwinian species : A group of organisms that shares an ancestor; a lineage that maintains its integrity with respect to other lineages through both time and space. At some point in the progress of such a group, members may diverge from one another: when such a divergence becomes sufficiently clear, the two populations are regarded as separate species. ; Microspecies : Species that reproduce without meiosis or mitosis so that each generation is genetically identical to the previous generation. See also apomixis. In practice, these definitions often coincide, and the differences between them are more a matter of emphasis than of outright contradiction. Nevertheless, no species concept yet proposed is entirely objective, or can be applied in all cases without resorting to judgement. Given the complexity of life, some have argued that such an objective definition is in all likelihood impossible, and biologists should settle for the most practical definition. For most vertebrates, this is the biological species concept, and to a lesser extent (or for different purposes) the phylogenetic species concept. Many BSC subspecies are considered species under the PSC; the difference between the BSC and the PSC can be summed up insofar as that the BSC defines a species as a consequence of manifest evolutionary history, while the PSC defines a species as a consequence of manifest evolutionary potential. Thus, a PSC species is "made" as soon as an evolutionary lineage has started to separate, while a BSC species starts to exist only when the lineage separation is complete.

Importance in biological classification

The idea of species has a long history. It is one of the most important levels of classification, for several reasons:
- It often corresponds to what lay people treat as the different basic kinds of organism - dogs are one species, cats another.
- It is the standard binomial nomenclature (or trinomial nomenclature) by which scientists typically refer to organisms.
- It is the only taxonomic level which has empirical content, in the sense that asserting that two animals are of different species is saying something more than classificatory about them. After thousands of years of use, the concept remains central to biology and a host of related fields, and yet also remains at times ill-defined and controversial.

Implications of assignment of species status

The naming of a particular species should be regarded as a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be confirmed or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as the same species. When two named species are discovered to be of the same species, the older species name is usually retained, and the newer species name dropped, a process called synonymization, or convivially, as lumping. Dividing a taxon into multiple, often new, taxons is called splitting. Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognizing differences or commonalities between organisms (see lumpers and splitters). Traditionally, researchers relied on observations of anatomical differences, and on observations of whether different populations were able to interbreed successfully, to distinguish species; both anatomy and breeding behavior are still important to assigning species status. As a result of the revolutionary (and still ongoing) advance in microbiological research techniques, including DNA analysis, in the last few decades, a great deal of additional knowledge about the differences and similarities between species has become available. Many populations which were formerly regarded as separate species are now considered to be a single taxon, and many formerly grouped populations have been split. Any taxonomic level (species, genus, family, etc.) can be synonymized or split, and at higher taxonomic levels, these revisions have been still more profound. From a taxonomical point of view, groups within a species can be defined as being of a taxon hierarchically lower than a species. In zoology only the subspecies is used, while in botany the variety, subvariety, and form are used as well.

The isolation species concept in more detail

In general, for large, complex, organisms that reproduce sexually (such as mammals and birds), one of several variations on the isolation or biological species concept is employed. Often, the distinction between different species, even quite closely related ones, is simple. Horses (Equus caballus) and donkeys (Equus asinus) are easily told apart even without study or training, and yet are so closely related that they can interbreed after a fashion. Because the result, a mule or hinny, is not usually fertile, they are clearly separate species. But many cases are more difficult to decide. This is where the isolation species concept diverges from the evolutionary species concept. Both agree that a species is a lineage that maintains its integrity over time, that is diagnosably different to other lineages (else we could not recognise it), is reproductively isolated (else the lineage would merge into others, given the chance to do so), and has a working intra-species recognition system (without which it could not continue). In practice, both also agree that a species must have its own independent evolutionary history—otherwise the characteristics just mentioned would not apply. The species concepts differ in that the evolutionary species concept does not make predictions about the future of the population: it simply records that which is already known. In contrast, the isolation species concept refuses to assign the rank of species to populations that, in the best judgement of the researcher, would recombine with other populations if given the chance to do so.

The isolation question

There are, essentially, two questions to resolve. First, is the proposed species consistently and reliably distinguishable from other species? Secondly, is it likely to remain so in the future? To take the second question first, there are several broad geographic possibilities.
- The proposed species are sympatric—they occupy the same habitat. Observation of many species over the years has failed to establish even a single instance of two diagnostically different populations that exist in sympatry and have then merged to form one united population. Without reproductive isolation, population differences cannot develop, and given reproductive isolation, gene flow between the populations cannot merge the differences. This is not to say that cross breeding does not take place at all, simply that it has become negligible. Generally, the hybrid individuals are less capable of successful breeding than pure-bred individuals of either species.
- The proposed species are allopatric—they occupy different geographical areas. Obviously, it is not possible to observe reproductive isolation in allopatric groups directly. Often it is not possible to achieve certainty by experimental means either: even if the two proposed species interbreed in captivity, this does not demonstrate that they would freely interbreed in the wild, nor does it always provide much information about the evolutionary fitness of hybrid individuals. A certain amount can be inferred from other experimental methods: for example, do the members of population A respond appropriately to playback of the recorded mating calls of population B? Sometimes, experiments can provide firm answers. For example, there are seven pairs of apparently almost identical marine snapping shrimp (Altheus) populations on either side of the Isthmus of Panama, which did not exist until about 3 million years ago. Until then, it is assumed, they were members of the same seven species. But when males and females from opposite sides of the isthmus are placed together, they fight instead of mating. Even if the isthmus were to sink under the waves again, the populations would remain genetically isolated: therefore they are now different species. In many cases, however, neither observation nor experiment can produce certain answers, and the determination of species rank must be made on a 'best guess' basis from a general knowledge of other related organisms.
- The proposed species are parapatric—they have breeding ranges that abut but do not overlap. This is fairly rare, particularly in temperate regions. The dividing line is often a sudden change in habitat (an ecotone) like the edge of a forest or the snow line on a mountain, but can sometimes be remarkably trivial. The parapatry itself indicates that the two populations occupy such similar ecological roles that they cannot coexist in the same area. Because they do not crossbreed, it is safe to assume that there is a mechanism, often behavioral, that is preventing gene flow between the populations, and that therefore they should be classified as separate species.
- There is a hybrid zone where the two populations mix. Typically, the hybrid zone will include representatives of one or both of the 'pure' populations, plus first-generation and back-crossing hybrids. The strength of the barrier to genetic transmission between the two pure groups can be assessed by the width of the hybrid zone relative to the typical dispersal distance of the organisms in question. The dispersal distance of oaks, for example, is the distance that a bird or squirrel can be expected to carry an acorn; the dispersal distance of Numbats is about 15 kilometres, as this is as far as young Numbats will normally travel in search of vacant territory to occupy after leaving the nest. The narrower the hybrid zone relative to the dispersal distance, the less gene flow there is between the population groups, and the more likely it is that they will continue on separate evolutionary paths. Nevertheless, it can be very difficult to predict the future course of a hybrid zone; the decision to define the two hybridizing populations as either the same species or as separate species is difficult and potentially controversial.
- The variation in the population is clinal; at either extreme of the population's geographic distribution, typical individuals are clearly different, but the transition between them is seamless and gradual. For example, the Koalas of northern Australia are clearly smaller and lighter in colour than those of the south, but there is no particular dividing line: the further south an individual Koala is found, the larger and darker it is likely to be; Koalas in intermediate regions are intermediate in weight and colour. In contrast, over the same geographic range, black-backed (northern) and white-backed (southern) Australian Magpies do not blend from one type to another: northern populations have black backs, southern populations white backs, and there is an extensive hybrid zone where both 'pure' types are common, as are crossbreeds. The variation in Koalas is clinal (a smooth transition from north to south, with populations in any given small area having a uniform appearance), but the variation in magpies is not clinal. In both cases, there is some uncertainty regarding correct classification, but the consensus view is that species rank is not justified in either. The gene flow between northern and southern magpie populations is judged to be sufficiently restricted to justify terming them subspecies (not full species); but the seamless way that local Koala populations blend one into another shows that there is substantial gene flow between north and south. As a result, experts tend to reject even subspecies rank in this case.

The difference question

Obviously, when defining a species, the geographic circumstances become meaningful only if the populations groups in question are clearly different: if they are not consistently and reliably distinguishable from one another, then we have no grounds for believing that they might be different species. The key question in this context, is "how different is different?" and the answer is usually "it all depends". In theory, it would be possible to recognise even the tiniest of differences as sufficient to delineate a separate species, provided only that the difference is clear and consistent (and that other criteria are met). There is no universal rule to state the smallest allowable difference between two species, but in general, very trivial differences are ignored on the twin grounds of simple practicality, and genetic similarity: if two population groups are so close that the distinction between them rests on an obscure and microscopic difference in morphology, or a single base substitution in a DNA sequence, then a demonstration of restricted gene flow between the populations will probably be difficult in any case. More typically, one or other of the following requirements must be met:
- It is possible to reliably measure a quantitative difference between the two groups that does not overlap. A population has, for example, thicker fur, rougher bark, longer ears, or larger seeds than another population, and although this characteristic may vary within each population, the two do not grade into one another, and given a reasonably large sample size, there is a definite discontinuity between them. Note that this applies to populations, not individual organisms, and that a small number of exceptional individuals within a population may 'break the rule' without invalidating it. The less a quantitative difference varies within a population and the more it varies between populations, the better the case for making a distinction. Nevertheless, borderline situations can only be resolved by making a 'best-guess' judgement.
- It is possible to distinguish a qualitative difference between the populations; a feature that does not vary continuously but is either entirely present or entirely absent. This might be a distinctively shaped seed pod, an extra primary feather, a particular courting behaviour, or a clearly different DNA sequence. Sometimes it is not possible to isolate a single difference between species, and several factors must be taken in combination. This is often the case with plants in particular. In eucalypts, for example, Corymbia ficifolia cannot be reliably distinguished from its close relative Corymbia calophylla by any single measure (and sometimes individual trees cannot be definitely assigned to either species), but populations of Corymbia can be clearly told apart by comparing the colour of flowers, bark, and buds, number of flowers for a given size of tree, and the shape of the leaves and fruit. When using a combination of characteristics to distinguish between populations, it is necessary to use a reasonably small number of factors (if more than a handful are needed, the genetic difference between the populations is likely to be insignificant and is unlikely to endure into the future), and to choose factors that are functionally independent (height and weight, for example, should usually be considered as one factor, not two).

Historical development of the species concept

In the earliest works of science, a species was simply an individual organism that represented a group of similar or nearly identical organisms. No other relationships beyond that group were implied. Aristotle used the words genus and species to mean generic and specific categories. Aristotle and other pre-Darwinian scientists took the species to be distinct and unchanging, with an "essence", like the chemical elements. When early observers began to develop systems of organization for living things, they began to place formerly isolated species into a context. To the modern mind, many of the schemes delineated are whimsical at best, such as those that determined consanguinity based on color (all plants with yellow flowers) or behavior (snakes, scorpions and certain biting ants). In the 18th century Carolus Linnaeus classified organisms according to differences in the form of reproductive apparatus. Although his system of classification sorts organisms according to degrees of similarity, it made no claims about the relationship between similar species. At the time, it was still widely believed that there is no organic connection between species, no matter how similar they appear; every species was individually created by God, a view today called creationism. This approach also suggested a type of idealism: the notion that each species exists as an "ideal form". Although there are always differences (although sometimes minute) between individual organisms, Linnaeus considered such variation problematic. He strove to identify individual organisms that were exemplary of the species, and considered other non-exemplary organisms to be deviant and imperfect. By the 19th century most naturalists understood that species could change form over time, and that the history of the planet provided enough time for major changes. As such, the new emphasis was on determining how a species could change over time. Jean-Baptiste Lamarck suggested that an organism could pass on an acquired trait to its offspring, i.e., the giraffe's long neck was attributed to generations of giraffes stretching to reach the leaves of higher treetops (this well-known and simplistic example, however, does not do justice to the breadth and subtlety of Lamarck's ideas). Lamarck's most important insight may have been that species can be extraordinarily fluid; his 1809 Zoological Philosophy contained one of the first logical refutations of creationism. With the acceptance of the work of Charles Darwin in the 1860s, Lamarck's view of evolution was quickly eclipsed. It was not until the late 20th century that his work began to be reexamined, and took its place as a fundamental stepping stone to the modern theory of adaptive mutation. Lamarck's long-discarded ideas of the goal-oriented evolution of species, also known the teleological process, have also received renewed attention, particularly by proponents of artificial selection. Charles Darwin and Alfred Wallace provided what scientists now consider the most powerful and compelling theory of evolution. Basically, Darwin argued that it is populations that evolve, not individuals. His argument relies on a radical shift in perspective from Linnaeus: rather than defining species in ideal terms (and searching for an ideal representative and rejecting deviations), Darwin considered variation among individuals to be natural. He further argued that variation, far from being problematic, actually provides the explanation for the existence of distinct species. Darwin's work drew on Thomas Malthus' insight that the rate of growth of a biological population will always outpace the rate of growth of the resources in the environment, such as the food supply. As a result, Darwin argued, not all the members of a population will be able to survive and reproduce. Those that did will, on average, be the ones possessing variations—however slight—that make them slightly better adapted to the environment. If these variable traits are heritable, then the offspring of the survivors will also possess them. Thus, over many generations, adaptive variations will accumulate in the population, while counter-adaptive will be eliminated. It should be emphasized that whether a variation is adaptive or non-adaptive depends on the environment: different environments favor different traits. Since the environment effectively selects which organisms live to reproduce, it is the environment (the "fight for existence") that selects the traits to be passed on. This is the theory of evolution by natural selection. In this model, the length of a giraffe's neck would be explained by positing that proto-giraffes with longer necks would have had a significant reproductive advantage to those with shorter necks. Over many generations, the entire population would be a species of long-necked animals. In 1859, when Darwin published his theory of natural selection, the mechanism behind the inheritance of individual traits was unknown. Although Darwin made some speculations on how traits are inherited (pangenesis), his theory relies only on the fact that inheritable traits exist, and are variable (which makes his accomplishment even more remarkable.) Although Gregor Mendel's paper on genetics was published in 1866, its significance was not recognized. It was not until 1900 that his work was rediscovered by Hugo de Vries, Carl Correns and Erich von Tschermak, who realised that the "inheritable traits" in Darwin's theory are genes. The theory of the evolution of species through natural selection has two important implications for discussions of species -- consequences that fundamentally challenge the assumptions behind Linnaeus' taxonomy. First, it suggests that species are not just similar, they may actually be related. Some students of Darwin argue that all species are descended from a common ancestor. Second, it supposes that "species" are not homogeneous, fixed, permanent things; members of a species are all different, and over time species change. This suggests that species do not have any clear boundaries but are rather momentary statistical effects of constantly changing gene-frequencies. One may still use Linnaeus' taxonomy to identify individual plants and animals, but one can no longer think of species as independent and immutable. The rise of a new species from a parental line is called speciation. There is no clear line demarcating the ancestral species from the descendant species. Although the current scientific understanding of species suggests there is no rigorous and comprehensive way to distinguish between different species in all cases, biologists continue to seek concrete ways to operationalize the idea. One of the most popular biological definitions of species is in terms of reproductive isolation; if two creatures cannot reproduce to produce fertile offspring, then they are in different species. This definition captures a number of intuitive species boundaries, but nonetheless has some problems, however. It has nothing to say about species that reproduce asexually, for example, and it is very difficult to apply to extinct species. Moreover, boundaries between species are often fuzzy: there are examples where members of one population can produce fertile offspring with a second population, and members of the second population can produce fertile offspring with members of a third population, but members of the first and third population cannot produces fertile offspring. Consequently, some people reject this notion of species. In recent years we have witnessed the drastic reduction in the size of breeding populations and the geographical range of many physically large mammals. In earlier times it was assumed that every species existed in at least a few thousand living individuals, except very rare relic, isolated groups. In the present, many well know mammal & bird species are so stressed by habitat loss, and other effects of the modern world, that only a very few breeding males may contribute the genetic material to a small number of breeding females. In these highly stressed conditions, the likelihood of change is very much greater. Mammals may become smaller, have darker fur, more stripes, more cautious behavior, even over time learn to co-exist with the human world. Very likely, evolution is radically accelerated, and we are only beginning to notice it. Species in transition before our eyes. It is possible that this severe stress is essential to the creation of new species, and may have been a prime factor throughout biological history, from other population reducing influences. Richard Dawkins defines two organisms as conspecific if and only if they have the same number of chromosomes and, for each chromosome, both organisms have the same number of nucleotides (The Blind Watchmaker, p. 118). However, most if not all taxonomists would strongly disagree. For example, in many amphibians, most notably in New Zealand's Leiopelma frogs, the genome consists of "core" chromosomes which are mostly invariable and accessory chromosomes, of which exist a number of possible combinations. Even though the chromosome numbers are highly variable between populations, these can interbreed successfully and form a single evolutionary unit. In plants, polyploidy is extremely commonplace with few restrictions on interbreeding; as individuals with an odd number of chromosome sets are usually sterile, depending on the actual number of chromosome sets present, this results in the odd situation where some individuals of the same evolutionary unit can interbreed with certain others and some cannot, with all populations being eventually linked as to form a common gene pool. The classification of species has been profoundly affected by technological advances that have allowed researchers to determine relatedness based on molecular markers, starting with the comparatively crude blood plasma precipitation assays in the mid-20th century and coming into full swing with Charles Sibley's ground-breaking DNA-DNA hybridisation studies in the 1970s. The results of the technique caused revolutionary changes in the higher taxonomic categories (such as phyla and classes), resulting in the reordering of many branches of the phylogenetic tree (see also: molecular phylogeny). For taxonomic categories below genera, the results have been mixed so far; the pace of evolutionary change on the molecular level is rather slow, yielding clear differences only after considerable periods of reproductive separation. Instances of hybridization can result in misleading molecular data, the Pomarine Skua - Great Skua phenomenon being a famous example. Turtles have been determined to evolve with just one-eighth of the speed of other reptiles on the molecular level, and the rate of molecular evolution in albatrosses is half of what is found in the rather closely related storm-petrels. The hybridization technique is however no longer considered a good technique and more reliable computational techniques for sequence comparison are now used for. Molecular taxonomy does not directly measure the evolutionary processes, but rather the overall change brought upon by these processes. The processes that lead to the generation and maintenance of variation such as mutation, crossover and selection are not uniform (see also molecular clock). DNA is only extremely rarely a direct target of natural selection rather than changes in the DNA sequence enduring over generations being a result of the latter; for example, silent transition-transversion combinations would alter the melting point of the DNA sequence, but not the sequence of the encoded proteins and thus are a possible example where, for example in microorganisms, a mutation confers a change in fitness all by itself.

External links

- http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/S/Speciation.html
- [http://www.sciencedaily.com/releases/2003/12/031231082553.htm 2003-12-31, ScienceDaily: Working On The 'Porsche Of Its Time': New Model For Species Determination Offered] Quote: "...two species of dinosaur that are members of the same genera varied from each other by just 2.2 percent. Translation of the percentage into an actual number results in an average of just three skeletal differences out of the total 338 bones in the body. Amazingly, 58 percent of these differences occurred in the skull alone. "This is a lot less variation than I'd expected", said Novak..."
- [http://www.sciencedaily.com/releases/2003/08/030808081854.htm 2003-08-08, ScienceDaily: Cross-species Mating May Be Evolutionarily Important And Lead To Rapid Change, Say Indiana University Researchers] Quote: "...the sudden mixing of closely related species may occasionally provide the energy to impel rapid evolutionary change..."
- [http://www.sciencedaily.com/releases/2004/01/040109064407.htm 2004-01-09 ScienceDaily: Mayo Researchers Observe Genetic Fusion Of Human, Animal Cells; May Help Explain Origin Of AIDS] Quote: "...The researchers have discovered conditions in which pig cells and human cells can fuse together in the body to yield hybrid cells that contain genetic material from both species... "What we found was completely unexpected", says Jeffrey Platt, M.D."
- [http://www.sciencedaily.com/releases/2000/09/000913211733.htm 2000-09-18, ScienceDaily: Scientists Unravel Ancient Evolutionary History Of Photosynthesis] Quote: "...gene-swapping was common among ancient bacteria early in evolution..."
- [http://plato.stanford.edu/entries/species/ Stanford Encyclopedia of Philosophy entry]
- [http://www.barcodinglife.org/ Barcoding of species] rank22 rank22 ms:Spesies ja:種 (生物) th:สปีชีส์

Pueraria

- Pueraria lobata
- Pueraria mirifica Pueraria is a genus of plants native to Southeastern Asia. Plants that belong to this genus include:
- Pueraria lobata (Kudzu)
- Pueraria mirifica Category:Faboideae

Fabaceae

Faboideae
Caesalpinioideae
Mimosoideae

References

[http://pgrc3.agr.ca/cgi-bin/npgs/html/family.pl?440 GRIN-CA] 2002-09-01 The Family Fabaceae sensu latu (also as Family Leguminosae) is a grouping of plants in the Order Fabales, and one of the largest families of flowering plants with 650 genera and over 18,000 species. These plants are commonly called legumes or pulses and the family contains some of our most valuable food crops, such as beans, peas, peanuts, soybeans, and lentils. Other members of the family are important sources of animal feed or green manure, such as lupins, clover, alfalfa, cassia, and soybean. Some genera such as Laburnum, Robinia, Gleditsia, Acacia, Mimosa, and Delonix are ornamental trees and shrubs. Still other members of the family have medicinal or insecticidal properties (for instance Derris) or yield important substances like gum arabic, tannin, dyes, or resins. Then there is kudzu, an east Asian species originally planted in the U.S. southeast for soil improvement and as a cattle feed, that has become a notorious invasive weed that tends to grow over everything. The use of the name Fabaceae is ambiguous, since it was used to represent the group faboideae for many years. Many specialists, especially in herbaria and floristics research, still use the old name of the family, Leguminosae, on order to avoid confusion. All members of this family have five-petaled flowers in which the superior ovary ripens to form a "pod", technically called a legume, whose two sides split apart, releasing the seeds which are attached to one or both seams. The legumous plants are classified into three subfamilies, sometimes raised to the rank of family in the order Fabales, on the basis of flower morphology (specifically, petal shape):
- Faboideae (Fabaceae sensu strictu), previously called Papilionoideae: One petal is large and has a crease in it, the two adjacent petals are on the sides, and the two bottom petals are joined together at the bottom, forming a boat-like structure.
- Caesalpinioideae (Caesalpiniaceae): The flowers are zygomorphic, but are very variable, e.g. closely resembling Faboideae flowers in Cercis, while symmetrical with five equal petals in Bauhinia.
- Mimosoideae (Mimosaceae): The petals are small, and are frequently globose or spicate and the stamens are the most showy part of the flower.

Nitrogen fixation

A significant characteristic of legumes is that they host bacteria in their roots, within structures called root nodules. These bacteria known as rhizobia have the ability to take nitrogen gas (N₂) out of the air and convert it to a form of nitrogen that is usable to the host plant ( NO₃^- or NH₃ ). This process is called nitrogen fixation. The legume, acting as a host; and rhizobia, acting as a provider of usable nitrate, form a symbiotic relationship. symbiotic

External link

- [http://www.ildis.org/ International Legume Database & Information Service] Category:Fabaceae Category:Legumes Category:Plant families Category:Nitrogen metabolism ja:マメ科

China

to protect the north from nomadic invaders and has been rebuilt several times since.]] China () refers to a number of states and cultures that have existed and are viewed as having succeeded one another in continental East Asia, dating back at least 3,500 years. China as it exists today has been variously described in different points of view as a single civilization or multiple civilizations, as a single state or multiple states, and as a single nation or multiple nations. With one of the world's longest periods of mostly uninterrupted civilization and the world's longest continuously used written language system, China's history has been largely characterized by repeated divisions and reunifications amid alternating periods of peace and war, and violent imperial dynastic change. The country's territorial extent expanded outwards from a core area in the North China Plain, and varied according to its moving fortunes to include multiple regions of East, Northeast, and Central Asia. For centuries, Imperial China was also one of the world's most technologically advanced civilizations, and East Asia's dominant cultural influence, with an impact lasting to the present day throughout the region. By the nineteenth and early twentieth centuries, however, China's political, economic, and military influence declined relative to growing regional power Japan and the influence of Western powers. Semi-colonialism developed by the late nineteenth century in parts of China, and the country was invaded by the Empire of Japan during World War II. The imperial system in China ended with the establishment of the Republic of China (ROC) under Sun Yat-sen in 1912; however, the next four decades of ROC rule were marred by warlord control, the Second Sino-Japanese War (WWII), and the Chinese Civil War which pitted Chinese Nationalists against the Communist forces. After its victory in the Chinese Civil War, the Communist Party of China under Mao Zedong established the People's Republic of China (PRC) in 1949, forcing the Republic of China (ROC) to retreat to the island of Taiwan, which it had governed since the end of World War II. Since then, the ROC has maintained administrative control over Taiwan, the Pescadores, several islands off the coast of Fujian province, and some islands in the South China Sea.

Terminology

"Zhongguo"

South China Sea China is called Zhongguo in Mandarin Chinese (Simplified: 中国, Traditional: 中國; also romanized as Jhongguo or Chung-kuo), which is usually translated as "Middle Kingdom", but could also be translated as "Central State" or "Central Country". Zhong (中) means "middle" or "center" while guo (国 or 國) means "country," "kingdom," "state," or "land", referring to the claim that China stood at the centre of that society's "known world", surrounded by lesser tributary states. The term has not been used consistently throughout Chinese history, however, and carries certain cultural and political connotations both positive and negative, some ideological, and early states considered part of Chinese history are not called "Zhongguo". During the Spring and Autumn Period, it was used only to describe the states politically descended from the Western Zhou Dynasty, in the Yellow River (Huang He) valley, to the exclusion of states such as Chu and Qin. The "Chinese" thus defined their nation as culturally and politically distinct from - and as the axis mundi of surrounding nations; a concept that continued well into the Qing Dynasty, although being continually redefined while the central political influence expanded territorially, and its culture assimilated alien influences. Thus Zhongguo quickly came to include areas farther south, as the cultural and political unit (not yet a "nation" or "country" in the modern sense) spread in a southerly direction, including the Yangtze River and Pearl River systems, and by the Tang Dynasty it even included "barbarian" regimes such as the Xianbei and Xiongnu. Inner Mongolia, Xinjiang, and Tibet, and the island of Taiwan, over time, came to be dominated (to a greater or lesser extent) by, or officially ruled by, imperial China, and are often included as a part of Zhongguo, though acceptance or denial of such claims remains politically controversial, especially where Zhongguo means PRC. During the Han Dynasty and before, Zhongguo had three distinctive meanings: # The area around the capital or imperial domain. The Book of Poetry explicitly gives this definition. # Territories under the direct authority of the "central" authorities. The Historical Records states: "Eight mountains are famed in the empire. Three are with the Man and Yi barbarians. Five are in Zhongguo." # The area now called the North China Plain. The Sanguo Zhi records the following monologue: "If we can lead the host of Wu and Yue (the area of southern Jiangsu and northern Zhejiang) to oppose Zhongguo, then we should break off relations with them soon." In this sense, the term is synonymous with Hua (華) and Xia (夏). During the period of division after the fall of the Han Dynasty, the term Zhongguo was subjected to transformation as a result of the surge of nomadic peoples from the northern frontier. This was doubly so after the loss of the Yellow River valley, the cradle of Chinese civilization, to these peoples. For example, the Xianbei called their Northern Wei regime Zhongguo, contrasting it with the Southern Dynasties, which they called the Yi (夷), meaning "barbarian". The southern dynasties, for their part, recently exiled from the north, called the Northern Wei Lu (虏), meaning "criminal" or "prisoner". In this way Zhongguo came to represent political legitimacy. It was used in this manner from the tenth century onwards by the competing dynasties of Liao, Jin and Song. The term Zhongguo came to be related to geographic, cultural and political identity and less to ethnic origin. The Republic of China, as it controlled mainland China, and later, the People's Republic of China, have used Zhongguo as an entity existing theoretically to mean all the territories and peoples within their political control as well as those outside of it (people in the Republic of China on Taiwan now usually use Zhongguo to refer to the PRC and use Taiwan to refer to itself). Thus it is asserted that all 56 officially recognized ethnic groups are Zhongguo ren (中國人), or Zhongguo people. Their disparate histories are collectively the history of Zhongguo.

"China"

Song in ancient times, was the imperial capital of 13 different historical dynasties (including the Han and Tang dynasties) in China.]] English and many other languages use forms of the name China (and the prefix Sino-), which is believed to have derived from the name of the Qin dynasty that first unified the country, even though it is not completely resolved and the origins are still controversial to an extent [http://www.bartleby.com/61/80/C0298000.html]. Despite the fact that the Qin dynasty was short-lived and was often regarded as overly tyrannical it unified the written language in China and gave the supreme ruler of China the title of "Emperor", hence, the subsequent Silk Road traders would identify themselves by that name. Alternate theories on the origin of the word "China" exist. In any circumstance, the word China passed through many languages along the Silk Road before it finally reached Europe and England. The Western "China", transliterated to Shina (支那) has also been used by Japanese since the nineteenth century, and has since evolved into a derogatory term in that language. The term "China" can narrowly mean China proper, or, often, China proper and Manchuria, Inner Mongolia, Tibet, and Xinjiang, a combination essentially coterminous with the 20th and 21st century political entity China; the boundaries between these regions do not necessarily follow provincial boundaries. In many contexts, "China" is commonly used to refer to the People's Republic of China or mainland China, while "Taiwan" is used to refer to the Republic of China. Informally, in economic or business contexts, "the Greater China region" (大中華地區) refers to Mainland China, Hong Kong, Macau, and Taiwan. Sinologists usually use "Chinese" in a more restricted sense, more akin to the classical usage of Zhongguo, or to the meaning of the "Han ethnic group", who make up the bulk of Mainland China. In many contexts it may be more appropriate to speak of "mainland China" (中國大陸,zhōngguó dàlù in Mandarin), especially when contrasting it with other, politically different regions like Hong Kong, Macau, and territories administered by the Republic of China (Taiwan).

History

:Main articles: History of China, History of the Republic of China (1912–1949; 1949–Present on Taiwan), History of People's Republic of China (1949–Present) History of People's Republic of China China was one of the earliest centers of human civilization. Chinese civilization was also one of the few to invent writing independently, the others being ancient Mesopotamia (Sumerians), India (Indus Valley Civilization), the Mayans, and, some hold, Ancient Egypt—though it may have been learned from the Sumerians. The first dynasty according to Chinese historical sources was the Xia Dynasty. Until scientific excavations were made at early bronze-age sites at Erlitou in Henan Province, it was difficult to separate myth from reality in regard to the existence of the Xia Dynasty. But since then, archaeologists have uncovered urban sites, bronze implements, and tombs that point to the possible existence of the Xia dynasty at the same locations cited in ancient Chinese historical texts. However, the first confirmed dynasty is the Shang, who settled along the Huang He river, dating from the 18th to the 12th centuries BC. The Shang were in turn invaded by the Zhou (12th to 5th centuries BC), whose centralized authority was slowly eroded by the ceding of state-like authority to warlords ruling small states; eventually, in the Spring and Autumn period, many strong independent states, in continuous war, paid but nominal deference to the Zhou state as the Imperial centre. They were all unified under one emperor in 221 BC by Qin Shi Huang, ushering in the Qin Dynasty, the first unified centralized Chinese state. This state, however, did not last for long, as it was way too authoritarian, destroying many sources of competition for power that were also sources of good governance and development, such as scholars and intellectuals. After the fall of authoritarian Qin Dynasty in 207 BC came the Han Dynasty which lasted until 220 AD. A period of disunion followed again. In 580, China was reunited under the Sui. Under the succeeding Tang and Song dynasties, China reached its golden age. For a long period of time, especially between the 7th and 14th centuries, China was one of the most advanced civilizations in the world in technology, literature, and art. The Song Dynasty fell to the invading Mongols in 1279. The Mongols, under Kublai Khan, established the Yuan Dynasty. A peasant named Zhu Yuanzhang overthrew the Mongols in 1368 and founded the Ming Dynasty, which lasted until 1644. After the Ming dynasty, came the Qing (Manchu) dynasty, which lasted until the overthrow of Puyi in 1911. Oftentimes regime change was violent and strongly opposed and the ruler class needed to take special measures to ensure their rule and the loyalty of the overthrown dynasty. For example, after the foreign Qing (Manchus) conquered China, because they were ever suspicious of the Han Chinese, the Qing rulers put into effect measures aimed at preventing the absorption of the Manchus into the dominant Han Chinese population. However, these restrictions proved ineffective against the assimilation of Manchus into the Chinese identity and culture. In the 18th century, China achieved a decisive technological advantage over the peoples of Central Asia, which it had been at war with for several centuries, while simultaneously falling behind Europe in that respect. This set the stage for the 19th century, in which China adopted a defensive posture against European imperialism while itself engaging in imperialistic expansion into Central Asia. See Imperialism in Asia. However the primary cause of the decline of the Chinese empire was not European and American interference, as the ethnocentric Western historians would lead many to believe. On the contrary it was a series of internal upheavals. Most prominent of these was the Taiping Civil War which lasted from 1851 to 1862. The civil war was started by an extremist believer in a school of thought partly influenced by Christianity who believed himself to be the son of God and the younger brother of Jesus. Although the imperial forces were eventually victorious, the civil war was one of the bloodiest in human history - costing at least twenty million lives (more than the total number of fatalities in the First World War). Prior to this conflict a number of Islamic Rebellions, especially in Central Asia, had occurred. Later, a second major rebellion took place, although this latter uprising was considerably smaller than the cataclysmic Taiping Civil War. This second conflict was the Boxer Rebellion which aimed to repel Westerners. Although secretly supporting the rebels, the Empress, Ci Xi, aided foreign forces in suppressing the uprising. Ci Xi, 1949.]] In 1912, after a prolonged period of decline, the institution of the Emperor of China disappeared and the Republic of China was established. The following three decades were a period of disunion — the Warlord Era, the Sino-Japanese War, and the Chinese Civil War. The latter ended in 1949 with the Communist Party of China in control of mainland China. The CPC established a communist state—the People's Republic of China—that laid claim to be the successor state of the Republic of China. Meanwhile, the disorganized and potentially corrupt ROC government of the Kuomintang fled to Taiwan, where it continued to be recognized as the legitimate government of all China by the Western bloc and the United Nations until the 1970s, when most nations and the UN switched recognition to the PRC. The United Kingdom and Portugal transferred their colonies of Hong Kong and Macau on the southern Chinese coast to the PRC in 1997 and 1999, respectively. China used in a modern context often refers to just the territory of the PRC, or to "Mainland China" (the territory of the PRC excluding Hong Kong and Macau). The PRC does not recognize the ROC, as it claims to have succeeded the ROC as the legitimate governing authority of all of China including Taiwan. On the other hand, the ROC—while never formally renouncing its earlier claims or changing official maps that show its territory as including both the modern-day PRC, Mongolia and Tibet—has moved away from this former identity representing its rule over all of China, and increasingly identifies itself as Taiwan. The PRC has historically resisted the ROC's identification of itself as Taiwan, especially in light of the movement supported by residents of Taiwan and others who advocate Taiwan's identity as an independent political entity. Significant disputes persist as to the nature and extent of China, possible Chinese reunification and the political status of Taiwan.

Chinese Pre-history

Archeological evidence suggests that the earliest occupants in China date as long as 2.24 million to 250,000 years ago by an ancient human relative (hominin) known as Homo erectus. One particular cave in Zhoukoudian (now known as Peking) has fossilised evidence dating to 300,000 and 550,000 years old. Evidence of primitive stone tool technology and animal bones in association to H. erectus have been studied since the late 18th century to 19th century in various areas of Eastern Asia including Indonesia (in particular the Island of Java) and Malaysia. Originally it is thought that these early hominis first evolved in Africa during the Pleistocene. It is thought that human evolution first took place in Africa expanding 7 million years. By 2 million years ago the first wave of migration from the species in association with H. erectus settled into various areas in the Old World. Fully modern humans (homo sapiens) are believed to originally have evolved roughly 200,000 and 168,000 years ago in Ethiopia or Southern Africa (ei. Homo sapiens idaltu). By 100,000 to 50,000 years ago modern human beings settled in all parts of the Old world (including the New World, Americas 25,000 to 11,000 BCE). By less than 100,000 years ago all proto-human populations disappeared as modern humans took over or drove other human species into extinction. It remains a controversial subject to whether fully modern humans evolved from separate H. erectus populations (known as "multiregional") as some evidence in ancient bones show a transitional change from H. erectus to H. sapiens having archaic features. However it is now more widely accepted that all modern humans genetically share a direct ancestor, a female nicknamed "Mitochondrial Eve" from Eastern Africa 150,000 years BCE. This model is known as Mitochondrial Eve Hypothesis. The earliest evidence examples of fully modern humans in China come from Liujiang, China where a cranium dates 67,000 years BCE. Another is a partial skeleton from Minatogawa being just 18,000 years old.

Political history

Before unification by the Qin Dynasty in 221 BC, "China" did not exist as a coherent entity. The Chinese civilization consisted of a patchwork of several states, each ruled by a king (王), duke (公), marquess (侯), or earl (伯). Although there was a central king who held nominal power, and powerful hegemons sometimes held considerable influence, each state was ruled as an independent political entity. This is also the time of the beginnings of Confucian philosophy and that of many other philosophies that greatly influenced Chinese philosophy-political thought. This ended with the Qin Dynasty unification, during which the office of the emperor was set up, and a system of bureaucratic administration established. After the Qin, China experienced about 13 more dynasties, many of which continued the extensive system of kingdoms, dukedoms, earldoms, and marquisates. The territory varied with several expansions and contractions depending on the strength of each emperor and dynasty. However the emperor had ultimate, supreme, and unquestionable authority as the political and religious leader of China. The emperor also consulted civil and martial ministers, especially the prime minister. Political power sometimes fell into the hands of powerful officials, eunuchs, or imperial relatives, often at the expense of a child heriditary emperor. This happened especially since the emperor often was many layers of power removed from the outside world, making him susceptible to manipulation because his sources for information could manipulate that information causing him to make incorrect decisions, especially when their age at becoming emperor often had no bottom limit, with rule passing heriditarily but also given "in trust" to another relative. Political relations with dependencies (tributary kingdoms) were maintained by international marriages, military aids, treaties, and gifts. (see section "Geography, Political" below for examples), Luoyang, Chang'an (today's Xi'an), Nanjing, and Beijing are the four cities most commonly designated as capitals of China over the course of history. Chinese was the official language, though periods of Mongol and Manchu conquest saw the arrival of Mongol and Manchu as alternate official languages. On January 1, 1912, the Republic of China (ROC) was established, signaling the end of the Manchu-dominated Qing Empire. Sun Yat-sen of the Kuomintang (KMT or Nationalist Party), was proclaimed provisional president of the republic. However, Yuan Shikai, a former Qing general who had defected to the revolutionary cause, soon forced Sun to step aside and took the presidency for himself (formally it was a negotiation where Sun agreed to step aside for what was then perceived as a strong reformer, Yuan). Before long, Yuan attempted to have himself proclaimed emperor of a new dynasty; however, he died soon of natural causes before fully taking power over all of the Chinese empire. After Yuan's downfall, China was politically fragmented, with an internationally-recognized, but virtually powerless, national government seated in Beijing (thus failing to fit the definition of a state). Warlords in various regions exercised actual control over their respective territories. state In the late 1920s, the Kuomintang, under Chiang Kai-shek, was able to reunify the country under its own control, moving the nation's capital to Nanjing and implementing "political tutelage", an intermediate stage of political development outlined in Sun Yat-sen's program for transforming China into a modern, democratic state. Effectively, political tutelage meant one-party rule by the Kuomintang with heavy Leninist influences. Ironically, both the Kuomintang and the CCP have heavy Leninist influences. In 1947, constitutional rule was established, but because of the ongoing Chinese Civil War between the Kuomintang and the Communist Party of China (CPC), many provisions of the 1947 ROC constitution were never put into actual practice on the mainland. By early 1950, the CPC had defeated the Kuomintang on the mainland, and the ROC government retreated to the island of Taiwan. Beginning in the late 1970s, Taiwan began the implementation of full, multi-party, representative democracy in the territories still under ROC control (i.e., Taiwan Province, Taipei, Kaohsiung and some offshore islands of Fujian province). Today, the political scene in the ROC is vibrant, with active participation by all sectors of society. But rather than the usual conservative-liberal policy distinctions that are the hallmarks of most democracies around the world, the main cleavage in ROC politics is the unification with China in the long-run vs. formal independence issue. However, Greens are generally more liberal (i.e. more environmentally friendly) and Blues are generally regarded as more conservative. environmentally friendly Meanwhile, Mao Zedong, the leader of the communists, proclaimed the People's Republic of China (PRC) on October 1, 1949 in Beijing, saying China had stood up. From the beginning, the PRC has been a dictatorial one-party state under the Communist Party. However, post-1978 reforms have led to the relaxation, in varying degrees, of party control over many areas of society. Nonetheless, the Communist Party still has absolute control over political aspects of society, and it continuously seeks to eradicate threats to its rule. Examples of this include the jailing of political opponents and journalists, general control of the press, regulation of religions and other non-party organizations, censorship of the press, literature and film, and suppression of independence/secessionist movements. In 1989, a popular demonstration held in Beijing at Tiananmen Square was violently put to an end by the Chinese government. Tiananmen_Square_protests_of_1989 The attempted eradication of the Falun Gong movement is also held by its supporters to be motivated by fear of Falun Gong's growing influence. Today, however, there is much more freedom in intellectual thought in non-political areas and propaganda, while still continuing, has lessened.

Territory

Historical overview

propaganda The Zhou Dynasty, which preceded the unification of China by Shi Huangdi, was originally the region around the Yellow River. Since then, the territory has expanded outward in all directions, and was largest during the Tang, Yuan, and Qing dynasties. The Qing Dynasty included parts of modern Russian Far East and Central Asia (west of Xinjiang). Xinjiang Along with provincial administrators, some foreign monarchs sent envoys to offer gifts to the Emperor of China and the Emperor returned compliments to them. The Chinese thought that the barbarians attached themselves to the virtue of the Emperor, while the foreign governments sometimes disagreed. Since the end of the 19th century, China has tried to reinterpret this relationship as suzerainty or suzerainty-dependency, but this no longer has any real conception in modern international political theories. The Qing Empire reduced the territorial value of the Great Wall of China as a barrier of China proper after they merged their homeland (Manchuria) north of the wall with China proper south of it. In 1683 after the surrender of the Kingdom of Tungning established by Koxinga, Taiwan including the Pescadores became a part of the Qing Empire, originally as one prefecture, then two, and later a province. Taiwan was subsequently ceded to Japan after the first Sino-Japanese War in 1895. At the end of the second Sino-Japanese War in 1945, Japan relinquished the sovereignty of the island in the San Francisco Peace Treaty, and the Republic of China took over. Since then, the de jure sovereignty of Taiwan has been under dispute between the PRC, and the now democratic ROC and Taiwan independence supporters.

Historical political divisions

Historically, top-level political divisions of China have altered as the administration changed. Top levels included circuits and provinces. Below that, there have been prefectures, subprefectures, departments, commanderies, districts, and counties. Recent divisions also include prefecture-level cities, county-level cities, towns and townships (see below for examples). Historically, most Chinese dynasties were based in the historical heartlands of China, known by the politically-correct term of China proper (since it doesn't include places it doesn't control, such as Mongolia or Taiwan). Various dynasties also exhibited expansionism by engaging in incursions into more peripheral territories like Inner Mongolia, Manchuria, Xinjiang, and Tibet. The Manchu-established Qing Dynasty and its successors, the Republic of China and the People's Republic of China cemented the incorporation of these territories into China. These territories are separated by borders that are vague at best, and do not correspond well to contemporary political divisions. China proper is generally thought to be bounded by the Great Wall and the edge of the Tibetan plateau; Manchuria and Inner Mongolia are found to the north of the Great Wall of China, and the boundary between them can either be taken as the present border between Inner Mongolia and the northeast Chinese provinces, or the more historic border of the World War II-era puppet state of Manchukuo; Xinjiang's borders correspond to today's administrative Xinjiang; and historic Tibet is conceived as occupying all of the Tibetan Plateau. China is also traditionally thought of as comprising North China (北方) and South China (南方), the geographic boundary between which north and south is largely generalized as Huai River (淮河) and Qinling Mountains (秦岭).

Geography and climate

Qinling Mountains

China within East Asia. ([http://www.cia.gov/cia/publications/factbook/reference_maps/pdf/asia.pdf PDF])

China is composed of a vast variety of highly different landscapes, with mostly plateaus and mountains in the west, and lower lands on the east. As a result, principal rivers flow from west to east, including the Yangtze (central), the Huang He (central-east), and the Amur (northeast), and sometimes toward the south (including the Pearl River, Mekong River, and Brahmaputra), with most Chinese rivers emptying into the Pacific. Most of China's arable lands lie along the two major rivers, the Yangtze and the Huang He, and each are the centers around which are founded China's major ancient civilizations. In the east, along the shores of the Yellow Sea and the East China Sea are found extensive and densely populated alluvial plains; the shore of the South China Sea is more mountainous and southern China is dominated by hill country and lower mountain ranges. To the west, the north has a great alluvial plain, and the south has a vast calcareous tableland traversed by hill ranges of moderate elevation, with the Himalayas, containing the highest point Mount Everest. The northwest also has high plateaus among more arid desert landscapes such as the Takla-Makan and the Gobi Desert, which has been expanding. Due to a prolonged drought and perhaps poor agricultural practices, dust storms have become usual in the spring in China. Dust blows all the way to southern China, Taiwan, and has even been measured on the West Coast of the United States. United States native to the bamboo forests of central and southern China.]] During many dynasties, the southwestern border of China has been the high mountains and deep valleys of Yunnan, which separate modern China from Burma, Laos and Vietnam. The climate of China varies greatly. The northern zone (within which lies Beijing) has a climate with winters of Arctic severity. The central zone (within which Shanghai is situated) has a generally temperate climate. The southern zone (within which lies Guangzhou and other southern provinces) has a generally subtropical climate. The Palaeozoic formations of China, excepting only the upper part of the Carboniferous system, are marine, while the Mesozoic and Tertiary deposits are estuarine and freshwater or else of terrestrial origin. Groups of volcanic cones occur in the Great Plain of north China. In the Liaodong and Shandong Peninsulas, there are basaltic plateaux.

Demographics

Shandong.]] Over a hundred ethnic groups have existed in China. In terms of numbers, however, the pre-eminent ethnic group in China is the Han, which is a group so diverse in its culture and language that some conceive of it as a larger overarching group bringing together many smaller, distinct ethnic groups sharing common traits in language and culture. Throughout history, many ethnic groups have been assimilated into neighbouring ethnicities or disappeared without a trace. Several previously distinct ethnic groups have been Sinicized into the Han, causing its population to increase dramatically; at the same time, many within the Han identity have maintained distinct linguistic and cultural traditions, though still identifying as Han. Many times in the past millenia many foreign groups have, in turn, shaped Han language and culture, for example the queue is a pig tail hairstyle strictly enforced by the Manchurians on the Han populace. The term Zhonghua Minzu is sometimes used to describe a notion of a "Chinese nationality" transcending ethnic divisions. The government of the People's Republic of China now officially recognizes a total of 56 ethnic groups, of which the largest is the Han Chinese. China's overall population is 1.3 billion. With the global human population currently estimated at about 6.4 billion, China is home to approximately 20%, or one-fifth of the human species, homo sapiens. The lack of birth control and promotion of population growth during the rule of Mao Zedong resulted in a demographic explosion, culminating in over 1.3 billion people today. As a response to the problems this is causing, the government of the PRC has enacted a birth control policy, commonly known as the One-child policy. The Han speak several mutually unintelligible tongues, classified by modern linguists as being separate languages, but regarded within the Chinese languages as "dialects" or "local languages" (topolects) within a single Chinese language (the word for "area languages" has an implication of dialect rather than a separate language, although on the basis of use, these topolects can be found to be separate and mutually unintelligible, and are so classified by many linguists). The various spoken varieties of Chinese share a common written standard, "Vernacular Chinese" or "baihua", which has been used since the early 20th Century and is based on Standard Mandarin, the standard spoken language, in grammar and vocabulary. In addition, another, more ancient written standard, Classical Chinese, was used for writing Chinese by the literati for thousands of years before the 20th Century. Classical Chinese is no longer the predominant form of written Chinese, though it continues to be a part of high school curricula and is hence intelligible to some degree to many Chinese people. Other than Standard Mandarin, spoken variants are usually not written; the exception is Standard Cantonese, which is sometimes written as Written Cantonese in informal contexts. Written Cantonese.]]

Culture

Religion

The major religions of China are:
- Taoism - exact numbers unknown
- Buddhism - exact numbers unknown [about 8%]
- Christianity - 2 to 4% (this is a Western number, the Chinese official number is much smaller than 1%)
- Islam - 1% to 2%
- Falun Gong - exact numbers unknown (claim not to be a "religion", though from a scholarly perspective is a spiritual practice, claimed numbers of followers of the Falun Dafa are also regarded as unreliable) While the People's Republic of China is officially atheist it does allow religion under strict supervision. Historically, Taoism and Buddhism has been the dominant religion of Chinese societies, and continues to be so in Chinese societies outside of direct PRC control. In recent years, Falun Gong, a spiritual practice drawing upon Buddhism and Taoism, has attracted great controversy after the government of the People's Republic of China labeled it an evil cult and began an attempt to eradicate it. The Falun Gong itself denies that it is a cult or a religion, even though there is solid evidence that determines Falun Gong as a rather" abormal" cult, several members have been seen to burn themselves alive even before the Chinese government has reacted to Falun Gong, unfortuantely, most people are oblivious of this fact and even a majority of members are oblivious to this. The Falun Gong says that it has approximately 70-100 million followers, which is a bit higher than estimates by outside groups, though exact numbers are unknown. They regularly protest against their suppression, both domestically and internationally.

Arts, scholarship, and literature

Falun Gong.]] Chinese literature has a long and prolific continuous history, in part because of the development of printmaking during the Song Dynasty. Before that, manuscripts of the Classics and religious texts (mainly Confucian, Taoist, and Buddhist) were manually written by ink brush (previously scratching shells) and distributed. Academies of scholars sponsored by the empire were formed to comment on these works in both printed and written form. Members of royalty frequently participated in these discussions. Tens of thousands of ancient written documents are still extant and more, from oracle bones to Qing edicts, are discovered each day, which had been formally ground up for use in Chinese medicine. oracle bones For centuries, opportunity for economic and social advancement in China could be provided by high performance on the imperial examinations. This led to a meritocracy, though in practice this was possible only among those who were not female or too poor to afford test preparation, as doing well still required tutorship. Nevertheless it was a system distinct from the European system of blood nobility. Imperial examinations required applicants to write essays and demonstrate mastery of the Confucian classics. Those who passed the highest level of the exam became elite scholar-officials known as jinshi, a highly esteemed socio-economic position. Chinese philosophers, writers, and poets have been, for the most part, highly respected, and played a key role in preserving and promoting the culture of the empire. Some classical scholars, however, were noted for their daring depictions of lives of the common people, often to the displeasure of authorities. (See List of Chinese authors, and List of Chinese language poets). The Chinese have created numerous musical instruments, such as the zheng, xiao, and erhu, that have spread throughout East and Southeast Asia, and especially areas under its influence. The sheng is the basis for several Western free-reed instruments. Chinese characters have had many variants and styles throughout the Chinese history, and were "simplified" in the mid-20th century on mainland China. Calligraphy is a major art-form in China, above that of painting and music. Because of its association with elite scholar-official bosses, it later on became commercialized, where works by famous artists became prized possessions. The great variation and beauty in the Chinese landscape is often the inspiration for great works of Chinese art. See Chinese painting for more details. Calligraphy, sushi, and bonsai are all millennia-old art that later spread to Japan and Korea.

Science and technology

Korea In addition to the cultural innovations mentioned above, technological inventions from China include:
- Compass
- Block Printmaking / Printing Technology
- Paper
- Asian abacus
- Gunpowder
- Crossbow
- Stirrup
- Lacquer
- Rudder
- Seismograph
- Silk
- Porcelain
- Paper money
- The Glider
- The Hot air balloon
- Fireworks
- Parachute Other areas of technological study:
- The main applications of mathematics in traditional China were architecture and geography. Pi (π) was calculated by 5th century mathematician Zu Chongzhi to the seventh digit. The decimal system was used in China as early as 14 Century BC. "Pascal's" Triangle was discovered by mathematician Liu Ju-Hsieh, long before Pascal was born.
- Studies in biology have been extensive, and historic records are consulted even today, such as pharmacopoeias of medicinal plant<